9S3D: PDB entry 9S3D
NAC bound human RNC with 58 amino acid ARF1-linker. Determined by electron microscopy at 2.32 Å resolution. Released 18 Feb 2026.
- Method
- Electron microscopy
- Resolution
- 2.32 Å
- Organism
- Homo sapiens
- Chains
- 87
- Atoms
- 218,816
- Mol. weight
- 4158.48 kDa
- Ligands
- ZN, MG
- Released
- 18 Feb 2026
Explore 9S3D in 3D
Show helices and sheets
RCSB PDB
PDBe
Secondary structure: helices and β-sheets
9S3D contains 566 α-helices and 579 β-strands across 81 chains. Residue ranges use author residue numbering, as in the PDB file, from PDBe. To see them in 3D, choose the Cartoon representation with Secondary structure coloring: helices, sheets and coils get different colors.
Chain CR: 15 helices, 19 β-strands
| Element | Residues | Length | Sheet |
|---|
| α-helix | 8-26 | 19 | |
| β-strand | 34-39 | 6 | 32 |
| α-helix | 45-59 | 15 | |
| α-helix | 65-82 | 18 | |
| β-strand | 93-101 | 9 | 32 |
| β-strand | 107-114 | 8 | 32 |
| β-strand | 124-128 | 5 | 32 |
| α-helix | 134-141 | 8 | |
| β-strand | 145-151 | 7 | 33 |
| β-strand | 155-162 | 8 | 33 |
| β-strand | 165-173 | 9 | 33 |
| α-helix | 176-179 | 4 | |
| α-helix | 184-186 | 3 | |
| α-helix | 187-212 | 26 | |
| β-strand | 214-215 | 2 | 34 |
| β-strand | 218-219 | 2 | 34 |
| β-strand | 224-228 | 5 | 33 |
| α-helix | 232-238 | 7 | |
| α-helix | 244-249 | 6 | |
| β-strand | 250-255 | 6 | 33 |
| α-helix | 261-296 | 36 | |
| β-strand | 301-303 | 3 | 35 |
| α-helix | 306-314 | 9 | |
| β-strand | 317-323 | 7 | 35 |
| β-strand | 329 | 1 | 36 |
| β-strand | 333-334 | 2 | 37 |
| β-strand | 341-342 | 2 | 37 |
| β-strand | 346 | 1 | 36 |
| α-helix | 348-353 | 6 | |
| α-helix | 364-368 | 5 | |
| α-helix | 374-385 | 12 | |
| β-strand | 388-392 | 5 | 35 |
| α-helix | 397-406 | 10 | |
| β-strand | 409-413 | 5 | 35 |
Chain CZ: 4 helices, 0 β-strands
| Element | Residues | Length | Sheet |
|---|
| α-helix | 64-66 | 3 | |
| α-helix | 74-76 | 3 | |
| α-helix | 81-87 | 7 | |
| α-helix | 88-92 | 5 | |
Chain La: 8 helices, 7 β-strands
| Element | Residues | Length | Sheet |
|---|
| α-helix | 3-5 | 3 | |
| α-helix | 7-10 | 4 | |
| α-helix | 42-48 | 7 | |
| β-strand | 72-74 | 3 | 64 |
| α-helix | 75-81 | 7 | |
| α-helix | 84-92 | 9 | |
| β-strand | 97-98 | 2 | 146 |
| α-helix | 99 | 1 | |
| β-strand | 100-102 | 3 | 65 |
| α-helix | 103-106 | 4 | |
| β-strand | 110-112 | 3 | 64 |
| β-strand | 123-126 | 4 | 65 |
| β-strand | 128-129 | 2 | 64 |
| α-helix | 131-140 | 10 | |
| β-strand | 143-146 | 4 | 65 |
Chain LA: 11 helices, 20 β-strands
| Element | Residues | Length | Sheet |
|---|
| α-helix | 4-5 | 2 | |
| β-strand | 18 | 1 | 77 |
| α-helix | 30-32 | 3 | |
| α-helix | 34-38 | 5 | |
| β-strand | 41-50 | 10 | 78 |
| α-helix | 51 | 1 | |
| β-strand | 57-64 | 8 | 78 |
| β-strand | 71-77 | 7 | 78 |
| β-strand | 79 | 1 | 79 |
| β-strand | 83 | 1 | 80 |
| β-strand | 87-90 | 4 | 78 |
| β-strand | 101-103 | 3 | 78 |
| α-helix | 104-106 | 3 | |
| β-strand | 112-113 | 2 | 81 |
| β-strand | 116 | 1 | 82 |
| β-strand | 126 | 1 | 82 |
| β-strand | 134-140 | 7 | 81 |
| α-helix | 142-144 | 3 | |
| β-strand | 145-149 | 5 | 81 |
| β-strand | 155-159 | 5 | 81 |
| β-strand | 163-165 | 3 | 78 |
| β-strand | 166 | 1 | 81 |
| β-strand | 169 | 1 | 79 |
| α-helix | 174-176 | 3 | |
| α-helix | 182-190 | 9 | |
| β-strand | 193 | 1 | 77 |
| α-helix | 198-200 | 3 | |
| α-helix | 201-203 | 3 | |
| β-strand | 224-225 | 2 | 83 |
| α-helix | 229-230 | 2 | |
| β-strand | 237-238 | 2 | 83 |
Chain Lb: 5 helices, 0 β-strands
| Element | Residues | Length | Sheet |
|---|
| α-helix | 12-18 | 7 | |
| α-helix | 22-24 | 3 | |
| α-helix | 37-48 | 12 | |
| α-helix | 49-51 | 3 | |
| α-helix | 54-75 | 22 | |
Chain LB: 18 helices, 19 β-strands
| Element | Residues | Length | Sheet |
|---|
| α-helix | 9-10 | 2 | |
| α-helix | 14-16 | 3 | |
| α-helix | 20-22 | 3 | |
| α-helix | 32-35 | 4 | |
| α-helix | 40-41 | 2 | |
| β-strand | 42-43 | 2 | 86 |
| β-strand | 45-59 | 15 | 87 |
| β-strand | 71-80 | 10 | 87 |
| α-helix | 82-83 | 2 | |
| β-strand | 84-95 | 12 | 86 |
| β-strand | 98-106 | 9 | 86 |
| α-helix | 112-115 | 4 | |
| α-helix | 116-118 | 3 | |
| α-helix | 134-137 | 4 | |
| α-helix | 140-156 | 17 | |
| β-strand | 159-165 | 7 | 86 |
| α-helix | 168-170 | 3 | |
| α-helix | 178-179 | 2 | |
| β-strand | 180-186 | 7 | 86 |
| α-helix | 190-199 | 10 | |
| β-strand | 204-205 | 2 | 86 |
| α-helix | 207-209 | 3 | |
| β-strand | 216-222 | 7 | 87 |
| β-strand | 227-229 | 3 | 88 |
| α-helix | 231-235 | 5 | |
| α-helix | 237-241 | 5 | |
| β-strand | 270-272 | 3 | 88 |
| β-strand | 276-288 | 13 | 87 |
| β-strand | 292 | 1 | 89 |
| β-strand | 298 | 1 | 89 |
| α-helix | 300-303 | 4 | |
| β-strand | 320 | 1 | 90 |
| β-strand | 324 | 1 | 90 |
| β-strand | 330-333 | 4 | 87 |
| β-strand | 344-349 | 6 | 87 |
| β-strand | 365-368 | 4 | 87 |
| α-helix | 382-389 | 8 | |
Chain Lc: 4 helices, 4 β-strands
| Element | Residues | Length | Sheet |
|---|
| α-helix | 11-21 | 11 | |
| β-strand | 26-29 | 4 | 91 |
| α-helix | 31-40 | 10 | |
| β-strand | 45-48 | 4 | 91 |
| α-helix | 54-67 | 14 | |
| β-strand | 70-73 | 4 | 91 |
| α-helix | 78-85 | 8 | |
| β-strand | 93-98 | 6 | 91 |
Chain LC: 22 helices, 11 β-strands
| Element | Residues | Length | Sheet |
|---|
| β-strand | 7-11 | 5 | 95 |
| β-strand | 17-23 | 7 | 95 |
| α-helix | 24-25 | 2 | |
| α-helix | 26-29 | 4 | |
| α-helix | 34-45 | 12 | |
| α-helix | 49-51 | 3 | |
| β-strand | 64-66 | 3 | 96 |
| β-strand | 77-79 | 3 | 96 |
| β-strand | 90 | 1 | 96 |
| α-helix | 100-101 | 2 | |
| α-helix | 117-130 | 14 | |
| α-helix | 134-140 | 7 | |
| β-strand | 152-154 | 3 | 95 |
| α-helix | 156-160 | 5 | |
| α-helix | 164-173 | 10 | |
| α-helix | 177-185 | 9 | |
| β-strand | 188-189 | 2 | 97 |
| α-helix | 190 | 1 | |
| α-helix | 193-197 | 5 | |
| β-strand | 201-202 | 2 | 97 |
| β-strand | 208-211 | 4 | 95 |
| α-helix | 217-221 | 5 | |
| β-strand | 228-231 | 4 | 95 |
| α-helix | 237-240 | 4 | |
| β-strand | 250-253 | 4 | 95 |
| α-helix | 254-264 | 11 | |
| α-helix | 279-282 | 4 | |
| α-helix | 289-293 | 5 | |
| α-helix | 296-299 | 4 | |
| α-helix | 302-305 | 4 | |
| α-helix | 312-314 | 3 | |
| α-helix | 322-328 | 7 | |
| α-helix | 331-363 | 33 | |
73 more chain groups are not listed. Open the entry in the viewer and use the sequence panel to see them.
Molecules and chains
| Molecule | Chains | Type | Length | Organism | UniProt |
|---|
| Nascent polypeptide-associated complex subunit alpha | NA | protein | 215 | Homo sapiens | Q13765 (AlphaFold model) |
| Isoform 2 of Transcription factor BTF3 | NB | protein | 162 | Homo sapiens | P20290 (AlphaFold model) |
| 58 amino acid ARF1-linker - V5 peptide - hCMV staller mRNA | CM | RNA | 586 | Homo sapiens | |
| prolyl-tRNA | CP | RNA | 75 | Homo sapiens | |
| Eukaryotic peptide chain release factor subunit 1 | CR | protein | 437 | Homo sapiens | P62495 (AlphaFold model) |
| nascent chain | CZ | protein | 95 | Homo sapiens | |
| 28S rRNA | L5 | RNA | 5070 | Homo sapiens | |
| 5S rRNA | L7 | RNA | 121 | Homo sapiens | |
| 5.8S rRNA | L8 | RNA | 157 | Homo sapiens | |
| 60S ribosomal protein L8 | LA | protein | 257 | Homo sapiens | P62917 (AlphaFold model) |
| 60S ribosomal protein L3 | LB | protein | 403 | Homo sapiens | P39023 |
| 60S ribosomal protein L4 | LC | protein | 427 | Homo sapiens | P36578 |
75 more molecules are not listed.
Sequence of entity 1 (NA), FASTA
>9S3D_1 Nascent polypeptide-associated complex subunit alpha (chains NA)
MPGEATETVPATEQELPQPQAETGSGTESDSDESVPELEEQDSTQATTQQAQLAAAAEID
EEPVSKAKQSRSEKKARKAMSKLGLRQVTGVTRVTIRKSKNILFVITKPDVYKSPASDTY
IVFGEAKIEDLSQQAQLAAAEKFKVQGEAVSNIQENTQTPTVQEESEEEEVDETGVEVKD
IELVMSQANVSRAKAVRALKNNSNDIVNAIMELTM
Sequence of entity 2 (NB), FASTA
>9S3D_2 Isoform 2 of Transcription factor BTF3 (chains NB)
MKETIMNQEKLAKLQAQVRIGGKGTARRKKKVVHRTATADDKKLQFSLKKLGVNNISGIE
EVNMFTNQGTVIHFNNPKVQASLAANTFTITGHAETKQLTEMLPSILNQLGADSLTSLRR
LAEALPKQSVDGKAPLATGEDDDDEVPDLVENFDEASKNEAN
Sequence of entity 3 (CM), FASTA
>9S3D_3 58 amino acid ARF1-linker - V5 peptide - hCMV staller mRNA (chains CM)
GGGCUUGUUCUUUUUGCAGAAGCUCAGAAUAAACGCUCAACUUUGGGCCGCCACCAUGGG
CUCCGACUACAAGGAUCAUGAUGGGGACUAUAAGGAUCACGACAUUGACUACAAAGACGA
UGACGAUAAGUCAGCUAGCGGUGAGAACCUGUACUUCCAGGGCAACAGCUUCAGCAAGCC
CCGGAAGGGCCUGUUCGGCAAAAAAGAAAUGCGCAUCCUCAUGGUGGGCCUGGAUGCUGC
AGGGAAGACCACGAUCCUCUACAAGCUUAAGCUGGGUGAGAUCGUGACCACCAUUCCCAC
CAUAGGCUUCAACGUGGAAACCGUGGAGUACAAGGGCAAACCGAUUCCGAACCCGCUGCU
GGGCCUGGAUAGCACCAUGGAACCGCUGGUGCUGAGUGCGAAAAAACUGAGCAGCCUGCU
GACCUGCAAAUAUAUUCCUCCUUAAGUUACCACUAAACCAGCCUCAAGAACACCCGAAUG
GAGUCUCUAAGCUACAUAAUACCAACUUACACUUUACAAAAUGUUGUCCCCCAAAAUGUA
GCCAUUCGUAUCUGCUCCUAAUAAAAAGAAAGUUUCUUCACAUUCU
Sequence of entity 4 (CP), FASTA
>9S3D_4 prolyl-tRNA (chains CP)
GGCUCGUUGGUCUAGGGGUAUGAUUCUCGCUUAGGGUGCGAGAGGUCCCGGGUUCAAAUC
CCGGACGAGCCCCCA
Sequence of entity 5 (CR), FASTA
>9S3D_5 Eukaryotic peptide chain release factor subunit 1 (chains CR)
MADDPSAADRNVEIWKIKKLIKSLEAARGNGTSMISLIIPPKDQISRVAKMLADEFGTAS
NIKSRVNRLSVLGAITSVQQRLKLYNKVPPNGLVVYCGTIVTEEGKEKKVNIDFEPFKPI
NTSLYLCDNKFHTEALTALLSDDSKFGFIVIDGSGALFGTLQGNTREVLHKFTVDLPKKH
GRGGQSALRFARLRMEKRHNYVRKVAETAVQLFISGDKVNVAGLVLAGSADFKTELSQSD
MFDQRLQSKVLKLVDISYGGENGFNQAIELSTEVLSNVKFIQEKKLIGRYFDEISQDTGK
YCFGVEDTLKALEMGAVEILIVYENLDIMRYVLHCQGTEEEKILYLTPEQEKDKSHFTDK
ETGQEHELIESMPLLEWFANNYKKFGATLEIVTDKSQEGSQFVKGFGGIGGILRYRVDFQ
GMEYQGGDDEFFDLDDY
Sequence of entity 6 (CZ), FASTA
>9S3D_6 nascent chain (chains CZ)
QGNSFSKPRKGLFGKKEMRILMVGLDAAGKTTILYKLKLGEIVTTIPTIGFNVETVEYKG
KPIPNPLLGLDSTMEPLVLSAKKLSSLLTCKYIPP
Sequence of entity 7 (L5), FASTA
>9S3D_7 28S rRNA (chains L5)
CGCGACCUCAGAUCAGACGUGGCGACCCGCUGAAUUUAAGCAUAUUAGUCAGCGGAGGAG
AAGAAACUAACCAGGAUUCCCUCAGUAACGGCGAGUGAACAGGGAAGAGCCCAGCGCCGA
AUCCCCGCCCCGCGGCGGGGCGCGGGACAUGUGGCGUACGGAAGACCCGCUCCCCGGCGC
CGCUCGUGGGGGGCCCAAGUCCUUCUGAUCGAGGCCCAGCCCGUGGACGGUGUGAGGCCG
GUAGCGGCCCCCGGCGCGCCGGGCCCGGGUCUUCCCGGAGUCGGGUUGCUUGGGAAUGCA
GCCCAAAGCGGGUGGUAAACUCCAUCUAAGGCUAAAUACCGGCACGAGACCGAUAGUCAA
CAAGUACCGUAAGGGAAAGUUGAAAAGAACUUUGAAGAGAGAGUUCAAGAGGGCGUGAAA
CCGUUAAGAGGUAAACGGGUGGGGUCCGCGCAGUCCGCCCGGAGGAUUCAACCCGGCGGC
GGGUCCGGCCGUGUCGGCGGCCCGGCGGAUCUUUCCCGCCCCCCGUUCCUCCCGACCCCU
CCACCCGCCCUCCCUUCCCCCGCCGCCCCUCCUCCUCCUCCCCGGAGGGGGCGGGCUCCG
GCGGGUGCGGGGGUGGGCGGGCGGGGCCGGGGGUGGGGUCGGCGGGGGACCGUCCCCCGA
CCGGCGACCGGCCGCCGCCGGGCGCAUUUCCACCGCGGCGGUGCGCCGCGACCGGCUCCG
GGACGGCUGGGAAGGCCCGGCGGGGAAGGUGGCUCGGGGGGCCCCGUCCGUCCGUCCGUC
CGUCCUCCUCCUCCCCCGUCUCCGCCCCCCGGCCCCGCGUCCUCCCUCGGGAGGGCGCGC
GGGUCGGGGCGGCGGCGGCGGCGGCGGUGGCGGCGGCGGCGGCGGCGGCGGGACCGAAAC
CCCCCCCGAGUGUUACAGCCCCCCCGGCAGCAGCACUCGCCGAAUCCCGGGGCCGAGGGA
GCGAGACCCGUCGCCGCGCUCUCCCCCCUCCCGGCGCCCACCCCCGCGGGGAAUCCCCCG
CGAGGGGGGUCUCCCCCGCGGGGGCGCGCCGGCGUCUCCUCGUGGGGGGGCCGGGCCACC
CCUCCCACGGCGCGACCGCUCUCCCACCCCUCCUCCCCGCGCCCCCGCCCCGGCGACGGG
GGGGGUGCCGCGCGCGGGUCGGGGGGCGGGGCGGACUGUCCCCAGUGCGCCCCGGGCGGG
UCGCGCCGUCGGGCCCGGGGGAGGUUCUCUCGGGGCCACGCGCGCGUCCCCCGAAGAGGG
GGACGGCGGAGCGAGCGCACGGGGUCGGCGGCGACGUCGGCUACCCACCCGACCCGUCUU
GAAACACGGACCAAGGAGUCUAACACGUGCGCGAGUCGGGGGCUCGCACGAAAGCCGCCG
UGGCGCAAUGAAGGUGAAGGCCGGCGCGCUCGCCGGCCGAGGUGGGAUCCCGAGGCCUCU
CCAGUCCGCCGAGGGCGCACCACCGGCCCGUCUCGCCCGCCGCGCCGGGGAGGUGGAGCA
CGAGCGCACGUGUUAGGACCCGAAAGAUGGUGAACUAUGCCUGGGCAGGGCGAAGCCAGA
GGAAACUCUGGUGGAGGUCCGUAGCGGUCCUGACGUGCAAAUCGGUCGUCCGACCUGGGU
AUAGGGGCGAAAGACUAAUCGAACCAUCUAGUAGCUGGUUCCCUCCGAAGUUUCCCUCAG
GAUAGCUGGCGCUCUCGCAGACCCGACGCACCCCCGCCACGCAGUUUUAUCCGGUAAAGC
GAAUGAUUAGAGGUCUUGGGGCCGAAACGAUCUCAACCUAUUCUCAAACUUUAAAUGGGU
AAGAAGCCCGGCUCGCUGGCGUGGAGCCGGGCGUGGAAUGCGAGUGCCUAGUGGGCCACU
UUUGGUAAGCAGAACUGGCGCUGCGGGAUGAACCGAACGCCGGGUUAAGGCGCCCGAUGC
CGACGCUCAUCAGACCCCAGAAAAGGUGUUGGUUGAUAUAGACAGCAGGACGGUGGCCAU
GGAAGUCGGAAUCCGCUAAGGAGUGUGUAACAACUCACCUGCCGAAUCAACUAGCCCUGA
AAAUGGAUGGCGCUGGAGCGUCGGGCCCAUACCCGGCCGUCGCCGGCAGUCGAGAGUGGA
CGGGAGCGGCGGGGGCGGCGCGCGCGCGCGCGCGUGUGGUGUGCGUCGGAGGGCGGCGGC
GGCGGCGGCGGCGGGGGUGUGGGGUCCUUCCCCCGCCCCCCCCCCCACGCCUCCUCCCCU
CCUCCCGCCCACGCCCCGCUCCCCGCCCCCGGAGCCCCGCGGACGCUACGCCGCGACGAG
UAGGAGGGCCGCUGCGGUGAGCCUUGAAGCCUAGGGCGCGGGCCCGGGUGGAGCCGCCGC
AGGUGCAGAUCUUGGUGGUAGUAGCAAAUAUUCAAACGAGAACUUUGAAGGCCGAAGUGG
AGAAGGGUUCCAUGUGAACAGCAGUUGAACAUGGGUCAGUCGGUCCUGAGAGAUGGGCGA
GCGCCGUUCCGAAGGGACGGGCGAUGGCCUCCGUUGCCCUCGGCCGAUCGAAAGGGAGUC
GGGUUCAGAUCCCCGAAUCCGGAGUGGCGGAGAUGGGCGCCGCGAGGCGUCCAGUGCGGU
AACGCGACCGAUCCCGGAGAAGCCGGCGGGAGCCCCGGGGAGAGUUCUCUUUUCUUUGUG
AAGGGCAGGGCGCCCUGGAAUGGGUUCGCCCCGAGAGAGGGGCCCGUGCCUUGGAAAGCG
UCGCGGUUCCGGCGGCGUCCGGUGAGCUCUCGCUGGCCCUUGAAAAUCCGGGGGAGAGGG
UGUAAAUCUCGCGCCGGGCCGUACCCAUAUCCGCAGCAGGUCUCCAAGGUGAACAGCCUC
UGGCAUGUUGGAACAAUGUAGGUAAGGGAAGUCGGCAAGCCGGAUCCGUAACUUCGGGAU
AAGGAUUGGCUCUAAGGGCUGGGUCGGUCGGGCUGGGGCGCGAAGCGGGGCUGGGCGCGC
GCCGCGGCUGGACGAGGCGCCGCCGCCCCCCCCACGCCCGGGGCACCCCCCUCGCGGCCC
UCCCCCGCCCCACCCCGCGCGCGCCGCUCGCUCCCUCCCCGCCCCGCGCCCUCUCUCUCU
CUCUCUCCCCCGCUCCCCGUCCUCCCCCCUCCCCGGGGGAGCGCCGCGUGGGGGCGGCGG
CGGGGGGAGAAGGGUCGGGGCGGCAGGGGCCGGCGGCGGCCCGCCGCGGGGCCCCGGCGG
CGGGGGCACGGUCCCCCGCGAGGGGGGCCCGGGCACCCGGGGGGCCGGCGGCGGCGGCGA
CUCUGGACGCGAGCCGGGCCCUUCCCGUGGAUCGCCCCAGCUGCGGCGGGCGUCGCGGCC
GCCCCCGGGGAGCCCGGCGGGCGCCGGCGCGCCCCCCCCCCCACCCCACGUCUCGUCGCG
CGCGCGUCCGCUGGGGGCGGGGAGCGGUCGGGCGGCGGCGGUCGGCGGGCGGCGGGGCGG
GGCGGUUCGUCCCCCCGCCCUACCCCCCCGGCCCCGUCCGCCCCCCGUUCCCCCCUCCUC
CUCGGCGCGCGGCGGCGGCGGCGGCAGGCGGCGGAGGGGCCGCGGGCCGGUCCCCCCCGC
CGGGUCCGCCCCCGGGGCCGCGGUUCCGCGCGGCGCCUCGCCUCGGCCGGCGCCUAGCAG
CCGACUUAGAACUGGUGCGGACCAGGGGAAUCCGACUGUUUAAUUAAAACAAAGCAUCGC
GAAGGCCCGCGGCGGGUGUUGACGCGAUGUGAUUUCUGCCCAGUGCUCUGAAUGUCAAAG
UGAAGAAAUUCAAUGAAGCGCGGGUAAACGGCGGGAGUAACUAUGACUCUCUUAAGGUAG
CCAAAUGCCUCGUCAUCUAAUUAGUGACGCGCAUGAAUGGAUGAACGAGAUUCCCACUGU
CCCUACCUACUAUCCAGCGAAACCACAGCCAAGGGAACGGGCUUGGCGGAAUCAGCGGGG
AAAGAAGACCCUGUUGAGCUUGACUCUAGUCUGGCACGGUGAAGAGACAUGAGAGGUGUA
GAAUAAGUGGGAGGCCCCCGGCGCCCCCCCGGUGUCCCCGCGAGGGGCCCGGGGCGGGGU
CCGCCGGCCCUGCGGGCCGCCGGUGAAAUACCACUACUCUGAUCGUUUUUUCACUGACCC
GGUGAGGCGGGGGGGCGAGCCCCGAGGGGCUCUCGCUUCUGGCGCCAAGCGCCCGGCCGC
GCGCCGGCCGGGCGCGACCCGCUCCGGGGACAGUGCCAGGUGGGGAGUUUGACUGGGGCG
GUACACCUGUCAAACGGUAACGCAGGUGUCCUAAGGCGAGCUCAGGGAGGACAGAAACCU
CCCGUGGAGCAGAAGGGCAAAAGCUCGCUUGAUCUUGAUUUUCAGUACGAAUACAGACCG
UGAAAGCGGGGCCUCACGAUCCUUCUGACCUUUUGGGUUUUAAGCAGGAGGUGUCAGAAA
AGUUACCACAGGGAUAACUGGCUUGUGGCGGCCAAGCGUUCAUAGCGACGUCGCUUUUUG
AUCCUUCGAUGUCGGCUCUUCCUAUCAUUGUGAAGCAGAAUUCACCAAGCGUUGGAUUGU
UCACCCACUAAUAGGGAACGUGAGCUGGGUUUAGACCGUCGUGAGACAGGUUAGUUUUAC
CCUACUGAUGAUGUGUUGUUGCCAUGGUAAUCCUGCUCAGUACGAGAGGAACCGCAGGUU
CAGACAUUUGGUGUAUGUGCUUGGCUGAGGAGCCAAUGGGGCGAAGCUACCAUCUGUGGG
AUUAUGACUGAACGCCUCUAAGUCAGAAUCCCGCCCAGGCGGAACGAUACGGCAGCGCCG
CGGAGCCUCGGUUGGCCUCGGAUAGCCGGUCCCCCGCCUGUCCCCGCCGGCGGGCCGCCC
CCCCCCUCCACGCGCCCCGCGCGCGCGGGAGGGCGCGUGCCCCGCCGCGCGCCGGGACCG
GGGUCCGGUGCGGAGUGCCCUUCGUCCUGGGAAACGGGGCGCGGCCGGAGAGGCGGCCGC
CCCCUCGCCCGUCACGCACCGCACGUUCGUGGGGAACCUGGCGCUAAACCAUUCGUAGAC
GACCUGCUUCUGGGUCGGGG
Sequence of entity 8 (L7), FASTA
>9S3D_8 5S rRNA (chains L7)
GUCUACGGCCAUACCACCCUGAACGCGCCCGAUCUCGUCUGAUCUCGGAAGCUAAGCAGG
GUCGGGCCUGGUUAGUACUUGGAUGGGAGACCGCCUGGGAAUACCGGGUGCUGUAGGCUU
U
Sequence of entity 9 (L8), FASTA
>9S3D_9 5.8S rRNA (chains L8)
CGACUCUUAGCGGUGGAUCACUCGGCUCGUGCGUCGAUGAAGAACGCAGCUAGCUGCGAG
AAUUAAUGUGAAUUGCAGGACACAUUGAUCAUCGACACUUCGAACGCACUUGCGGCCCCG
GGUUCCUCCCGGGGCUACGCCUGUCUGAGCGUCGCUU
Sequence of entity 10 (LA), FASTA
>9S3D_10 60S ribosomal protein L8 (chains LA)
MGRVIRGQRKGAGSVFRAHVKHRKGAARLRAVDFAERHGYIKGIVKDIIHDPGRGAPLAK
VVFRDPYRFKKRTELFIAAEGIHTGQFVYCGKKAQLNIGNVLPVGTMPEGTIVCCLEEKP
GDRGKLARASGNYATVISHNPETKKTRVKLPSGSKKVISSANRAVVGVVAGGGRIDKPIL
KAGRAYHKYKAKRNCWPRVRGVAMNPVEHPFGGGNHQHIGKPSTIRRDAPAGRKVGLIAA
RRTGRLRGTKTVQEKEN
Sequence of entity 11 (LB), FASTA
>9S3D_11 60S ribosomal protein L3 (chains LB)
MSHRKFSAPRHGSLGFLPRKRSSRHRGKVKSFPKDDPSKPVHLTAFLGYKAGMTHIVREV
DRPGSKVNKKEVVEAVTIVETPPMVVVGIVGYVETPRGLRTFKTVFAEHISDECKRRFYK
NWHKSKKKAFTKYCKKWQDEDGKKQLEKDFSSMKKYCQVIRVIAHTQMRLLPLRQKKAHL
MEIQVNGGTVAEKLDWARERLEQQVPVNQVFGQDEMIDVIGVTKGKGYKGVTSRWHTKKL
PRKTHRGLRKVACIGAWHPARVAFSVARAGQKGYHHRTEINKKIYKIGQGYLIKDGKLIK
NNASTDYDLSDKSINPLGGFVHYGEVTNDFVMLKGCVVGTKKRVLTLRKSLLVQTKRRAL
EKIDLKFIDTTSKFGHGRFQTMEEKKAFMGPLKKDRIAKEEGA
Sequence of entity 12 (LC), FASTA
>9S3D_12 60S ribosomal protein L4 (chains LC)
MACARPLISVYSEKGESSGKNVTLPAVFKAPIRPDIVNFVHTNLRKNNRQPYAVSELAGH
QTSAESWGTGRAVARIPRVRGGGTHRSGQGAFGNMCRGGRMFAPTKTWRRWHRRVNTTQK
RYAICSALAASALPALVMSKGHRIEEVPELPLVVEDKVEGYKKTKEAVLLLKKLKAWNDI
KKVYASQRMRAGKGKMRNRRRIQRRGPCIIYNEDNGIIKAFRNIPGITLLNVSKLNILKL
APGGHVGRFCIWTESAFRKLDELYGTWRKAASLKSNYNLPMHKMINTDLSRILKSPEIQR
ALRAPRKKIHRRVLKKNPLKNLRIMLKLNPYAKTMRRNTILRQARNHKLRVDKAAAAAAA
LQAKSDEKAAVAGKKPVVGKKGKKAAVGVKKQKKPLVGKKAAATKKPAPEKKPAEKKPTT
EEKKPAA
Ligands and cofactors
| ID | Name | Formula | Copies |
|---|
| ZN | Zinc ion | Zn | 8 |
| MG | Magnesium ion | Mg | 191 |
Primary citation
Structural basis of co-translational N-myristoylation in humans. Denk, T., Monassa, P., Musial, J. et al. Nat Commun (2026) 17:1191-1191. DOI 10.1038/s41467-025-67962-4 · PubMed
Other PDB entries of the same protein (UniProt Q13765 (AlphaFold model), which also has an AlphaFold model), best resolution first:
- 3MCB 1.9 Å, Crystal structure of NAC domains of human nascent polypeptide-associated complex (NAC)
- 9I2D 2.19 Å, NMT1-NAC bound human RNC with 10 amino acid ARF1-linker
- 9S3B 2.38 Å, NMT1-NAC bound human RNC with 58 amino acid ARF1-linker - State 1
- 3MCE 2.4 Å, Crystal structure of the NAC domain of alpha subunit of nascent polypeptide-associated…
- 9S3C 2.42 Å, NMT1-NAC bound human RNC with 58 amino acid ARF1-linker - State 2
- 9QLO 2.47 Å, NMT1-NAC bound human RNC with full length ARF1 - State 1
- 3LKX 2.5 Å, Human nac dimerization domain
- 9QLQ 2.57 Å, NMT1-NAC bound human RNC with full length ARF1 - alternative State
- 9MR4 2.65 Å, NAC: Ribosome nascent chain complex(Oxa1L)
- 9T6D 2.67 Å, NAA40-NAC bound human 80S (combined translation states)
- 28LN 2.7 Å, XBP1u-stalled RPL4 RNC in complex with NAC
- 9RG7 2.72 Å, In vitro reconstituted NAA40-NAC bound 80S
Browse structure collections
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