11AA: Nucleolar protein 16

Yeast co-transcriptional pre-60S assembly intermediate Nop12 RNP. Determined by electron microscopy at 2.78 Å resolution. Released 1 Jul 2026.

Method
Electron microscopy
Resolution
2.78 Å
Organism
Saccharomyces cerevisiae S288C
Chains
17
Atoms
29,525
Mol. weight
1795.11 kDa
Ligands
MG
Released
1 Jul 2026

Explore 11AA in 3D Show helices and sheets RCSB PDB PDBe

Secondary structure: helices and β-sheets

11AA contains 165 α-helices and 111 β-strands across 14 chains. Residue ranges use author residue numbering, as in the PDB file, from PDBe. To see them in 3D, choose the Cartoon representation with Secondary structure coloring: helices, sheets and coils get different colors.

Chain 7: 3 helices, 0 β-strands

ElementResiduesLengthSheet
α-helix179-19214
α-helix196-2005
α-helix212-22514
Chain b: 8 helices, 16 β-strands
ElementResiduesLengthSheet
β-strand33-37517
β-strand38118
α-helix43-5513
β-strand59-61317
α-helix63-642
β-strand65118
α-helix73-808
β-strand85-92817
β-strand96-102717
β-strand109-1181017
β-strand130119
β-strand133120
α-helix134-1352
β-strand136-139417
α-helix141-1444
α-helix147-16014
α-helix162-1632
β-strand170120
β-strand175-182817
β-strand185-1951117
β-strand210-214517
β-strand218-2271017
β-strand235-238417
α-helix245-25410
Chain G: 16 helices, 6 β-strands
ElementResiduesLengthSheet
α-helix55-6814
β-strand70-7121
α-helix721
α-helix73-764
α-helix77-793
α-helix84-9411
α-helix95-973
α-helix98-1014
α-helix102-11817
α-helix127-1293
β-strand133-13422
α-helix136-1449
β-strand150-15452
α-helix161-17313
β-strand177-18042
α-helix183-1897
β-strand197-20042
α-helix205-2073
α-helix208-21811
α-helix219-2235
α-helix224-2263
β-strand234-23521
Chain H: 11 helices, 14 β-strands
ElementResiduesLengthSheet
α-helix143-1475
β-strand149-15683
α-helix158-1614
α-helix163-17210
β-strand203-20973
β-strand21314
α-helix219-2257
β-strand22814
β-strand234-24183
α-helix246-2538
β-strand25815
β-strand26315
β-strand265-26843
α-helix272-2743
α-helix277-2793
β-strand280-28456
α-helix292-2998
α-helix300-3023
β-strand305-31286
β-strand319-32796
α-helix330-3378
β-strand34317
β-strand36317
β-strand365-36846
α-helix369-3702
Chain i: 7 helices, 0 β-strands
ElementResiduesLengthSheet
α-helix22-243
α-helix28-303
α-helix32-343
α-helix35-4814
α-helix52-6312
α-helix66-7712
α-helix80-9819
Chain K: 18 helices, 13 β-strands
ElementResiduesLengthSheet
α-helix34-4815
β-strand64-6528
α-helix70-745
β-strand77-8489
α-helix961
β-strand97-101510
α-helix108-1147
α-helix120-1212
β-strand123-127511
α-helix130-1345
α-helix137-1448
β-strand153-156411
α-helix157-1626
α-helix167-1748
β-strand179-183511
α-helix187-1893
α-helix190-1934
α-helix198-2003
β-strand207-209311
β-strand212-213212
β-strand216-217212
α-helix219-23214
β-strand233-237510
β-strand243-25089
α-helix256-27318
β-strand276-28389
β-strand289-29359
α-helix296-3016
α-helix328-3369
α-helix339-3413
α-helix342-3454
Chain L: 6 helices, 6 β-strands
ElementResiduesLengthSheet
α-helix13-2513
α-helix52-532
β-strand54113
β-strand57-59314
α-helix63-664
β-strand69-71314
β-strand75115
α-helix77-826
α-helix87-937
β-strand95113
β-strand97115
α-helix106-12116
Chain m: 7 helices, 3 β-strands
ElementResiduesLengthSheet
β-strand196117
α-helix198-20811
α-helix212-2143
α-helix217-2193
β-strand222-224317
α-helix229-2324
α-helix239-26325
α-helix270-2712
β-strand278119
α-helix282-2909

6 more chain groups are not listed. Open the entry in the viewer and use the sequence panel to see them.

Molecules and chains

MoleculeChainsTypeLengthOrganismUniProt
25S ribosomal RNA1RNA3396Saccharomyces cerevisiae S288C
5.8S ribosomal RNA2RNA159Saccharomyces cerevisiae S288C
Internal transcribed spacer 2 ITS23RNA232Saccharomyces cerevisiae S288C
Nucleolar protein 167protein231Saccharomyces cerevisiae S288CP40007 (AlphaFold model)
60S ribosomal protein L8-AGprotein256Saccharomyces cerevisiae S288CP17076 (AlphaFold model)
Nucleolar protein 12Hprotein459Saccharomyces cerevisiae S288CQ08208 (AlphaFold model)
Proteasome-interacting protein CIC1Kprotein376Saccharomyces cerevisiae S288CP38779 (AlphaFold model)
60S ribosomal protein L13-ALprotein199Saccharomyces cerevisiae S288CQ12690
60S ribosomal protein L15-ANprotein204Saccharomyces cerevisiae S288CP05748
Ribosome biogenesis protein BRX1bprotein291Saccharomyces cerevisiae S288CQ08235
60S ribosomal protein L36-Aiprotein100Saccharomyces cerevisiae S288CP05745
rRNA-processing protein EBP2mprotein427Saccharomyces cerevisiae S288CP36049

5 more molecules are not listed.

Sequence of entity 1 (1), FASTA
>11AA_1 25S ribosomal RNA (chains 1)
GUUUGACCUCAAAUCAGGUAGGAGUACCCGCUGAACUUAAGCAUAUCAAUAAGCGGAGGA
AAAGAAACCAACCGGGAUUGCCUUAGUAACGGCGAGUGAAGCGGCAAAAGCUCAAAUUUG
AAAUCUGGUACCUUCGGUGCCCGAGUUGUAAUUUGGAGAGGGCAACUUUGGGGCCGUUCC
UUGUCUAUGUUCCUUGGAACAGGACGUCAUAGAGGGUGAGAAUCCCGUGUGGCGAGGAGU
GCGGUUCUUUGUAAAGUGCCUUCGAAGAGUCGAGUUGUUUGGGAAUGCAGCUCUAAGUGG
GUGGUAAAUUCCAUCUAAAGCUAAAUAUUGGCGAGAGACCGAUAGCGAACAAGUACAGUG
AUGGAAAGAUGAAAAGAACUUUGAAAAGAGAGUGAAAAAGUACGUGAAAUUGUUGAAAGG
GAAGGGCAUUUGAUCAGACAUGGUGUUUUGUGCCCUCUGCUCCUUGUGGGUAGGGGAAUC
UCGCAUUUCACUGGGCCAGCAUCAGUUUUGGUGGCAGGAUAAAUCCAUAGGAAUGUAGCU
UGCCUCGGUAAGUAUUAUAGCCUGUGGGAAUACUGCCAGCUGGGACUGAGGACUGCGACG
UAAGUCAAGGAUGCUGGCAUAAUGGUUAUAUGCCGCCCGUCUUGAAACACGGACCAAGGA
GUCUAACGUCUAUGCGAGUGUUUGGGUGUAAAACCCAUACGCGUAAUGAAAGUGAACGUA
GGUUGGGGCCUCGCAAGAGGUGCACAAUCGACCGAUCCUGAUGUCUUCGGAUGGAUUUGA
GUAAGAGCAUAGCUGUUGGGACCCGAAAGAUGGUGAACUAUGCCUGAAUAGGGUGAAGCC
AGAGGAAACUCUGGUGGAGGCUCGUAGCGGUUCUGACGUGCAAAUCGAUCGUCGAAUUUG
GGUAUAGGGGCGAAAGACUAAUCGAACCAUCUAGUAGCUGGUUCCUGCCGAAGUUUCCCU
CAGGAUAGCAGAAGCUCGUAUCAGUUUUAUGAGGUAAAGCGAAUGAUUAGAGGUUCCGGG
GUCGAAAUGACCUUGACCUAUUCUCAAACUUUAAAUAUGUAAGAAGUCCUUGUUACUUAA
UUGAACGUGGACAUUUGAAUGAAGAGCUUUUAGUGGGCCAUUUUUGGUAAGCAGAACUGG
CGAUGCGGGAUGAACCGAACGUAGAGUUAAGGUGCCGGAAUACACGCUCAUCAGACACCA
CAAAAGGUGUUAGUUCAUCUAGACAGCCGGACGGUGGCCAUGGAAGUCGGAAUCCGCUAA
GGAGUGUGUAACAACUCACCGGCCGAAUGAACUAGCCCUGAAAAUGGAUGGCGCUCAAGC
GUGUUACCUAUACUCUACCGUCAGGGUUGAUAUGAUGCCCUGACGAGUAGGCAGGCGUGG
AGGUCAGUGACGAAGCCUAGACCGUAAGGUCGGGUCGAACGGCCUCUAGUGCAGAUCUUG
GUGGUAGUAGCAAAUAUUCAAAUGAGAACUUUGAAGACUGAAGUGGGGAAAGGUUCCACG
UCAACAGCAGUUGGACGUGGGUUAGUCGAUCCUAAGAGAUGGGGAAGCUCCGUUUCAAAG
GCCUGAUUUUAUGCAGGCCACCAUCGAAAGGGAAUCCGGUUAAGAUUCCGGAACCUGGAU
AUGGAUUCUUCACGGUAACGUAACUGAAUGUGGAGACGUCGGCGCGAGCCCUGGGAGGAG
UUAUCUUUUCUUCUUAACAGCUUAUCACCCCGGAAUUGGUUUAUCCGGAGAUGGGGUCUU
AUGGCUGGAAGAGGCCAGCACCUUUGCUGGCUCCGGUGCGCUUGUGACGGCCCGUGAAAA
UCCACAGGAAGGAAUAGUUUUCAUGCCAGGUCGUACUGAUAACCGCAGCAGGUCUCCAAG
GUGAACAGCCUCUAGUUGAUAGAAUAAUGUAGAUAAGGGAAGUCGGCAAAAUAGAUCCGU
AACUUCGGGAUAAGGAUUGGCUCUAAGGGUCGGGUAGUGAGGGCCUUGGUCAGACGCAGC
GGGCGUGCUUGUGGACUGCUUGGUGGGGCUUGCUCUGCUAGGCGGACUACUUGCGUGCCU
UGUUGUAGACGGCCUUGGUAGGUCUCUUGUAGACCGUCGCUUGCUACAAUUAACGAUCAA
CUUAGAACUGGUACGGACAAGGGGAAUCUGACUGUCUAAUUAAAACAUAGCAUUGCGAUG
GUCAGAAAGUGAUGUUGACGCAAUGUGAUUUCUGCCCAGUGCUCUGAAUGUCAAAGUGAA
GAAAUUCAACCAAGCGCGGGUAAACGGCGGGAGUAACUAUGACUCUCUUAAGGUAGCCAA
AUGCCUCGUCAUCUAAUUAGUGACGCGCAUGAAUGGAUUAACGAGAUUCCCACUGUCCCU
AUCUACUAUCUAGCGAAACCACAGCCAAGGGAACGGGCUUGGCAGAAUCAGCGGGGAAAG
AAGACCCUGUUGAGCUUGACUCUAGUUUGACAUUGUGAAGAGACAUAGAGGGUGUAGAAU
AAGUGGGAGCUUCGGCGCCAGUGAAAUACCACUACCUUUAUAGUUUCUUUACUUAUUCAA
UGAAGCGGAGCUGGAAUUCAUUUUCCACGUUCUAGCAUUCAAGGUCCCAUUCGGGGCUGA
UCCGGGUUGAAGACAUUGUCAGGUGGGGAGUUUGGCUGGGGCGGCACAUCUGUUAAACGA
UAACGCAGAUGUCCUAAGGGGGGCUCAUGGAGAACAGAAAUCUCCAGUAGAACAAAAGGG
UAAAAGCCCCCUUGAUUUUGAUUUUCAGUGUGAAUACAAACCAUGAAAGUGUGGCCUAUC
GAUCCUUUAGUCCCUCGGAAUUUGAGGCUAGAGGUGCCAGAAAAGUUACCACAGGGAUAA
CUGGCUUGUGGCAGUCAAGCGUUCAUAGCGACAUUGCUUUUUGAUUCUUCGAUGUCGGCU
CUUCCUAUCAUACCGAAGCAGAAUUCGGUAAGCGUUGGAUUGUUCACCCACUAAUAGGGA
ACGUGAGCUGGGUUUAGACCGUCGUGAGACAGGUUAGUUUUACCCUACUGAUGAAUGUUA
CCGCAAUAGUAAUUGAACUUAGUACGAGAGGAACAGUUCAUUCGGAUAAUUGGUUUUUGC
GGCUGUCUGAUCAGGCAUUGCCGCGAAGCUACCAUCCGCUGGAUUAUGGCUGAACGCCUC
UAAGUCAGAAUCCAUGCUAGAACGCGGUGAUUUCUUUGCUCCACACAAUAUAGAUGGAUA
CGAAUAAGGCGUCCUUGUGGCGUCGCUGAACCAUAGCAGGCUAGCAACGGUGCACUUGGC
GGAAAGGCCUUGGGUGCUUGCUGGCGAAUUGCAAUGUCAUUUUGCGUGGGGAUAAAUCAU
UUGUAUACGACUUAGAUGUACAACGGGGUAUUGUAAGCAGUAGAGUAGCCUUGUUGUUAC
GAUCUGCUGAGAUUAAGCCUUUGUUGUCUGAUUUGU
Sequence of entity 2 (2), FASTA
>11AA_2 5.8S ribosomal RNA (chains 2)
AAAACUUUCAACAACGGAUCUCUUGGUUCUCGCAUCGAUGAAGAACGCAGCGAAAUGCGA
UACGUAAUGUGAAUUGCAGAAUUCCGUGAAUCAUCGAAUCUUUGAACGCACAUUGCGCCC
CUUGGUAUUCCAGGGGGCAUGCCUGUUUGAGCGUCAUUU
Sequence of entity 3 (3), FASTA
>11AA_3 Internal transcribed spacer 2 ITS2 (chains 3)
CCUUCUCAAACAUUCUGUUUGGUAGUGAGUGAUACUCUUUGGAGUUAACUUGAAAUUGCU
GGCCUUUUCAUUGGAUGUUUUUUUUCCAAAGAGAGGUUUCUCUGCGUGCUUGAGGUAUAA
UGCAAGUACGGUCGUUUUAGGUUUUACCAACUGCGGCUAAUCUUUUUUUAUACUGAGCGU
AUUGGAACGUUAUCGAUAAGAAGAGAGCGUCUAGGCGAACAAUGUUCUUAAA
Sequence of entity 4 (7), FASTA
>11AA_4 Nucleolar protein 16 (chains 7)
MTSVRKRKMNRSSVGKATRRNKDKQRKINIQSNPIIAANWDYSLTMAQNYKKLGLRAKLQ
TPAGGKEADLSKVVKRIPLTKPVLDEDEDEDEGEDEQNDYNAATVELDENEIPEGGARIQ
RDKNGDVVRVVYGKKKNFDADEDVNEIKARDTTEETEVVKKLEELASRPVIRKERSQSER
EEEWLEKLYKKHGDDYKKMFFDKKLNIYQQSEGDLKRRLLRWKKRNGIASK
Sequence of entity 5 (G), FASTA
>11AA_5 60S ribosomal protein L8-A (chains G)
MAPGKKVAPAPFGAKSTKSNKTRNPLTHSTPKNFGIGQAVQPKRNLSRYVKWPEYVRVQR
QKKILSIRLKVPPTIAQFQYTLDRNTAAETFKLFNKYRPETAAEKKERLTKEAAAVAEGK
SKQDASPKPYAVKYGLNHVVALIENKKAKLVLIANDVDPIELVVFLPALCKKMGVPYAIV
KGKARLGTLVNQKTSAVAALTEVRAEDEAALAKLVSTIDANFADKYDEVKKHWGGGILGN
KAQAKMDKRAKNSDSA
Sequence of entity 6 (H), FASTA
>11AA_6 Nucleolar protein 12 (chains H)
MSSAIDNLFGNIDEKKIESSVDKLFSSSCGPINKLEVKSKTRTVLPDSKKRERAAEADQE
EKEASKPDVSDEQTEEVALPKVKKAKKSKRNDEDEDLEARYYAKLLNEEAEAEDDKPTVT
KTDETSVPLTSAAKKVDFKEDELEKAERTVFIGNILSTVITSKKVYKEFKKLFGTNPIAE
TEESGNEKEEESSKKSDNNEFAIESIRFRSISFDEALPRKVAFVQQKFHKSRDTINAYIV
YKNKSAVRKICSNLNAVVFQDHHLRVDSVAHPAPHDKKRSIFVGNLDFEEIEESLWKHFE
PCGDIEYVRIIRDSKTNMGKGFAYVQFKDLQSVNKALLLNEKPMKSQKQEDENTKKPTKK
ARKLRVSRCKNMKKGTTIGTGLDRNGLTDSQRTRAGRAKKILGKADRATLGQEITIEGLR
AKKGEGSTHLKKKKQRSATGRVTKRSIAFKKAQAEKSKK
Sequence of entity 7 (K), FASTA
>11AA_7 Proteasome-interacting protein CIC1 (chains K)
MAKKSNSKKSTPVSTPSKEKKKVIEKKSSTAIPRERVIKAVNELIKFTSKPQDENNEEGN
NGKKNLLEDDEEELKKDLQLIVVNNKSFTGTSKSFKLKLLNVKHSFYKPWKEASATAVKD
FKVLLILKDSDIKKVSEDDLFDQLDSEGIKVDEIICGKDLKTVYKAYEARNAFISQFSLI
LADDSIVTSLPKLMGGKAYNKVETTPISIRTHANKEFSLTTLTNNIKKVYMNQLPVKLPR
GTTLNVHLGNLEWLRPEEFVDNVELISEQLIKAYQIRSIFIKTNRSPVLPLYYNQDVLDE
LEAKKDKIEETHEDDMVTIDGVQVHLSTFNKGLMEIANPSELGSIFSKQINNAKKRSSSE
LEKESSESEAVKKAKS
Sequence of entity 8 (L), FASTA
>11AA_8 60S ribosomal protein L13-A (chains L)
MAISKNLPILKNHFRKHWQERVKVHFDQAGKKVSRRNARATRAAKIAPRPLDLLRPVVRA
PTVKYNRKVRAGRGFTLAEVKAAGLTAAYARTIGIAVDHRRQNRNQEIFDANVQRLKEYQ
SKIIVFPRNGKAPEAEQVLSAAATFPIAQPATDVEARAVQDNGESAFRTLRLARSEKKFR
GIREKRAREKAEAEAEKKK
Sequence of entity 9 (N), FASTA
>11AA_9 60S ribosomal protein L15-A (chains N)
MGAYKYLEELQRKKQSDVLRFLQRVRVWEYRQKNVIHRAARPTRPDKARRLGYKAKQGFV
IYRVRVRRGNRKRPVPKGATYGKPTNQGVNELKYQRSLRATAEERVGRRAANLRVLNSYW
VNQDSTYKYFEVILVDPQHKAIRRDARYNWICDPVHKHREARGLTATGKKSRGINKGHKF
NNTKAGRRKTWKRQNTLSLWRYRK
Sequence of entity 10 (b), FASTA
>11AA_10 Ribosome biogenesis protein BRX1 (chains b)
MSSIYKALAGKSKDNKSEKKQGNVKQFMNKQRTLLISSRGVNYRHRHLIQDLSGLLPHSR
KEPKLDTKKDLQQLNEIAELYNCNNVLFFEARKHQDLYLWLSKPPNGPTIKFYIQNLHTM
DELNFTGNCLKGSRPVLSFDQRFESSPHYQLIKELLVHNFGVPPNARKSKPFIDHVMSFS
IVDDKIWVRTYEISHSTKNKEEYEDGEEDISLVEIGPRFVMTVILILEGSFGGPKIYENK
QYVSPNVVRAQIKQQAAEEAKSRAEAAVERKIKRRENVLAADPLSNDALFK
Sequence of entity 11 (i), FASTA
>11AA_11 60S ribosomal protein L36-A (chains i)
MTVKTGIAIGLNKGKKVTSMTPAPKISYKKGAASNRTKFVRSLVREIAGLSPYERRLIDL
IRNSGEKRARKVAKKRLGSFTRAKAKVEEMNNIIAASRRH
Sequence of entity 12 (m), FASTA
>11AA_12 rRNA-processing protein EBP2 (chains m)
MAKGFKLKELLSHQKEIEKAEKLENDLKKKKSQELKKEEPTIVTASNLKKLEKKEKKADV
KKEVAADTEEYQSQALSKKEKRKLKKELKKMQEQDATEAQKHMSGDEDESGDDREEEEEE
EEEEEGRLDLEKLAKSDSESEDDSESENDSEEDEDVVAKEESEEKEEQEEEQDVPLSDVE
FDSDADVVPHHKLTVNNTKAMKHALERVQLPWKKHSFQEHQSVTSETNTDEHIKDIYDDT
ERELAFYKQSLDAVLVARDELKRLKVPFKRPLDYFAEMVKSDEHMDKIKGKLIEEASDKK
AREEARRQRQLKKFGKQVQNATLQKRQLEKRETLEKIKSLKNKRKHNEIDHSEFNVGVEE
EVEGKRFDRGRPNGKRAAKNAKYGQGGMKRFKRKNDATSSADVSGFSSRKMKGKTNRPGK
SRRARRF

Ligands and cofactors

IDNameFormulaCopies
MGMagnesium ionMg5

Primary citation

Mechanism for the initiation of co-transcriptional pre-60 S assembly. Piwowarczyk, R., Klinge, S. bioRxiv (2026). DOI 10.64898/2026.05.22.727207 · PubMed

Other PDB entries of the same protein (UniProt P40007 (AlphaFold model), which also has an AlphaFold model), best resolution first:

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