6PSV: Protein TraR

Escherichia coli RNA polymerase promoter unwinding intermediate (TpreRPo) with TraR and rpsT P2 promoter. Determined by electron microscopy at 3.5 Å resolution. Released 25 Mar 2020.

Method
Electron microscopy
Resolution
3.5 Å
Organism
Escherichia coli
Chains
10
Atoms
32,331
Mol. weight
562.9 kDa
Ligands
ZN, 1N7, MG
Released
25 Mar 2020

Explore 6PSV in 3D Show helices and sheets RCSB PDB PDBe

Secondary structure: helices and β-sheets

6PSV contains 174 α-helices and 227 β-strands across 8 chains. Residue ranges use author residue numbering, as in the PDB file, from PDBe. To see them in 3D, choose the Cartoon representation with Secondary structure coloring: helices, sheets and coils get different colors.

Chain G: 6 helices, 19 β-strands

ElementResiduesLengthSheet
β-strand13-1752
β-strand23-3192
α-helix35-4915
β-strand5313
β-strand56-5944
β-strand6114
β-strand7415
α-helix78-803
α-helix82-854
β-strand90-9126
β-strand97-10484
β-strand108-11145
α-helix112-1143
β-strand115-11624
β-strand122-12326
β-strand129-13355
β-strand140-14784
β-strand14913
β-strand15317
α-helix155-1584
β-strand171-17224
β-strand17517
β-strand180-189102
β-strand199-20792
α-helix213-23321
Chain H: 6 helices, 19 β-strands
ElementResiduesLengthSheet
β-strand1918
β-strand23-3198
α-helix321
α-helix35-4915
β-strand54-6189
β-strand74110
α-helix78-869
β-strand90111
β-strand99-10139
β-strand104-105212
β-strand109-111313
α-helix112-1143
β-strand123111
β-strand129-132413
β-strand133110
β-strand139-140212
β-strand142-14879
β-strand152-153214
β-strand17219
β-strand175-176214
β-strand180-18348
β-strand188-19038
β-strand198-207108
α-helix213-22715
α-helix228-2303
Chain I: 62 helices, 102 β-strands
ElementResiduesLengthSheet
α-helix5-73
β-strand13-14215
α-helix19-202
α-helix29-3810
α-helix49-568
α-helix571
β-strand59-60216
β-strand66-751016
α-helix83-886
β-strand93-1051316
α-helix1061
α-helix1131
β-strand114-118516
β-strand121-128816
α-helix1291
β-strand130117
β-strand136-137217
β-strand142-145417
α-helix1461
β-strand147-151518
α-helix1521
β-strand154-158519
β-strand173-177519
β-strand180120
β-strand186119
β-strand187-188221
β-strand194-196321
β-strand204-205221
α-helix207-2126
α-helix217-2248
β-strand227-231522
β-strand238-240322
α-helix244-2463
β-strand250123
β-strand255-257324
β-strand260-263424
β-strand268123
α-helix271-2766
α-helix277-2815
β-strand284-286322
α-helix289-2924
β-strand296-297222
β-strand315-316222
α-helix319-3268
β-strand332-336522
β-strand339125
β-strand343125
α-helix346-3538
α-helix359-37012
α-helix378-39013
β-strand396120
α-helix399-40810
α-helix422-43716
α-helix442-4443
β-strand451-454418
α-helix456-48025
α-helix496-50712
β-strand512-514317
α-helix515-5162
α-helix521-5244
β-strand529126
β-strand530-531218
β-strand535127
β-strand537127
α-helix552-5543
β-strand558128
β-strand574126
β-strand575128
β-strand576129
β-strand580-581230
β-strand587-588230
β-strand589-594631
β-strand598132
β-strand599-606831
α-helix608-6136
β-strand616131
β-strand623133
β-strand628132
β-strand629133
β-strand633-636434
β-strand637131
β-strand643-646434
α-helix647-6493
β-strand652-654331
β-strand662129
α-helix676-68510
α-helix686-6883
β-strand689135
β-strand700115
α-helix705-7106
β-strand716-717236
β-strand722-727637
β-strand731-736637
α-helix738-7403
β-strand749-752437
β-strand757-758238
β-strand763139
β-strand764-765238
β-strand768-769236
β-strand776-777237
β-strand782-785436
β-strand789140
β-strand794140
β-strand798-804741
β-strand816-819441
α-helix821-8244
β-strand830-831242
β-strand832139
β-strand834-841843
β-strand842-843244
β-strand846-847244
β-strand849145
α-helix859-8624
β-strand865146
β-strand870147
β-strand871146
α-helix8721
β-strand882-884347
β-strand887-888245
α-helix897-9026
α-helix903-9075
α-helix909-9113
β-strand914-915245
β-strand918-919247
α-helix920-9212
β-strand927-935943
α-helix940-9423
α-helix943-98038
α-helix986-9894
α-helix994-9974
α-helix1007-103630
β-strand1046-10551043
β-strand1057-1058242
β-strand1066-1067248
β-strand1076-1080541
β-strand1087-1088249
α-helix10921
β-strand1093149
α-helix10941
β-strand1096141
α-helix1102-11054
α-helix1110-112920
α-helix1139-115012
α-helix1166-117510
β-strand1182-1183215
α-helix1186-11883
α-helix1192-120110
β-strand1209-1210241
β-strand1212-1213249
α-helix12191
β-strand1220149
α-helix12211
β-strand1225-1232841
β-strand1233-1234248
β-strand1235135
α-helix1239-12424
β-strand1244-1246350
β-strand1252151
β-strand1268-1269252
α-helix1273-12808
α-helix1284-12907
α-helix1298-130912
α-helix1321-133111
β-strand1335-1337353
Chain J: 60 helices, 81 β-strands
ElementResiduesLengthSheet
β-strand22-24353
α-helix27-337
β-strand34154
β-strand44155
β-strand51155
α-helix521
α-helix59-624
β-strand84156
β-strand91156
α-helix95-995
β-strand102154
β-strand103-1121057
α-helix123-1264
α-helix132-1398
β-strand144-145258
β-strand159-160258
α-helix162-17110
β-strand179158
α-helix181-19010
α-helix195-20713
α-helix213-22917
α-helix234-2374
β-strand238-244757
β-strand252-254359
α-helix2551
β-strand260-262359
α-helix265-28521
α-helix289-30719
β-strand317160
β-strand323160
α-helix3241
α-helix3371
α-helix338-3425
β-strand346-347252
β-strand350-354550
β-strand355-356261
β-strand365-369561
α-helix370-3767
α-helix378-38811
α-helix394-40310
α-helix406-41510
β-strand421-424450
α-helix431-4333
β-strand434-437450
β-strand438-442561
β-strand447-448261
α-helix454-4574
β-strand465-469550
α-helix474-4796
α-helix480-4845
β-strand485150
α-helix486-4883
β-strand491162
β-strand498162
α-helix505-5139
β-strand526-527263
α-helix530-5378
β-strand547-5561063
β-strand564-5731063
α-helix574-5807
α-helix589-5913
α-helix598-61215
α-helix615-63319
α-helix645-6473
α-helix650-67021
α-helix675-70228
β-strand706-707264
β-strand714-715264
α-helix721-7277
β-strand746-748365
α-helix7491
β-strand754-758565
α-helix769-78618
α-helix790-80314
β-strand809-811366
β-strand821-822267
β-strand826168
β-strand831168
α-helix835-8395
β-strand843169
β-strand844170
α-helix8471
β-strand848-849271
α-helix8501
β-strand856-858371
β-strand863169
α-helix866-8749
β-strand880-881267
β-strand882170
β-strand894-895266
α-helix896-8994
β-strand901172
β-strand908172
α-helix909-9102
α-helix915-92410
α-helix925-9284
β-strand956-961673
β-strand965-967374
β-strand973-975374
β-strand981-985573
β-strand991-996673
β-strand1002-1003274
β-strand1010-1011273
β-strand1019174
β-strand1033-1035375
β-strand1038176
β-strand1042177
β-strand1046-1048377
β-strand1059-1061377
β-strand1077-1079376
β-strand1080-1081275
β-strand1087175
β-strand1089-1090278
β-strand1093-1096478
β-strand1098-1100376
β-strand1106-1107277
β-strand1114-1115275
β-strand1122-1123277
α-helix1139-11457
α-helix1153-11553
β-strand1156179
β-strand1162-1165480
β-strand1170181
β-strand1172181
β-strand1173-1178680
β-strand1186-1190580
α-helix1204-12052
β-strand1209179
α-helix1217-12193
α-helix1220-12245
α-helix1226-124419
α-helix1251-126010
β-strand1263-1266482
β-strand1279-1281382
α-helix1282-129413
β-strand1302-1304382
α-helix1311-13144
α-helix1320-13256
α-helix1328-133710
α-helix1347-13526
α-helix1361-137212
Chain K: 4 helices, 0 β-strands
ElementResiduesLengthSheet
α-helix9-124
α-helix18-3114
α-helix46-5510
α-helix62-7514
Chain L: 26 helices, 4 β-strands
ElementResiduesLengthSheet
α-helix97-10610
α-helix115-13521
α-helix138-15215
α-helix158-1603
β-strand162183
α-helix214-23219
α-helix245-25511
α-helix256-2583
β-strand261183
α-helix263-28826
α-helix299-3068
α-helix327-3293
α-helix334-35118
α-helix355-39137
α-helix401-41717
α-helix421-4233
α-helix427-44519
α-helix454-47421
α-helix480-4867
α-helix493-5008
α-helix502-5043
β-strand505-506259
α-helix519-5224
β-strand524151
α-helix529-5302
α-helix531-55020
α-helix553-56210
α-helix573-5775
α-helix584-59916
α-helix605-6106
Chain M: 5 helices, 0 β-strands
ElementResiduesLengthSheet
α-helix251-2533
α-helix264-27310
α-helix278-2814
α-helix286-2905
α-helix297-30812
Chain N: 5 helices, 2 β-strands
ElementResiduesLengthSheet
α-helix4-2522
β-strand3611
α-helix421
β-strand4311
α-helix441
α-helix46-516
α-helix59-7012

Molecules and chains

MoleculeChainsTypeLengthOrganismUniProt
Protein TraRNprotein72Escherichia coliP41065 (AlphaFold model)
DNA-directed RNA polymerase subunit alphaG, H, Mprotein329Escherichia coliP0A7Z4 (AlphaFold model)
DNA-directed RNA polymerase subunit betaIprotein1342Escherichia coliP0A8V2 (AlphaFold model)
DNA-directed RNA polymerase subunit beta'Jprotein1430Escherichia coliP0A8T7 (AlphaFold model)
DNA-directed RNA polymerase subunit omegaKprotein91Escherichia coliP0A800
RNA polymerase sigma factor RpoDLprotein616Escherichia coliP00579
DNA (85-mer)ODNA85Escherichia coli
DNA (85-mer)PDNA85Escherichia coli
Sequence of entity 1 (N), FASTA
>6PSV_1 Protein TraR (chains N)
SDEADEAYSVTEQLTMTGINRIRQKINAHGIPVYLCEACGNPIPEARRKIFPGVTLCVEC
QAYQERQRKHYA
Sequence of entity 2 (G, H, M), FASTA
>6PSV_2 DNA-directed RNA polymerase subunit alpha (chains G, H, M)
MQGSVTEFLKPRLVDIEQVSSTHAKVTLEPLERGFGHTLGNALRRILLSSMPGCAVTEVE
IDGVLHEYSTKEGVQEDILEILLNLKGLAVRVQGKDEVILTLNKSGIGPVTAADITHDGD
VEIVKPQHVICHLTDENASISMRIKVQRGRGYVPASTRIHSEEDERPIGRLLVDACYSPV
ERIAYNVEAARVEQRTDLDKLVIEMETNGTIDPEEAIRRAATILAEQLEAFVDLRDVRQP
EVKEEKPEFDPILLRPVDDLELTVRSANCLKAEAIHYIGDLVQRTEVELLKTPNLGKKSL
TEIKDVLASRGLSLGMRLENWPPASIADE
Sequence of entity 3 (I), FASTA
>6PSV_3 DNA-directed RNA polymerase subunit beta (chains I)
MVYSYTEKKRIRKDFGKRPQVLDVPYLLSIQLDSFQKFIEQDPEGQYGLEAAFRSVFPIQ
SYSGNSELQYVSYRLGEPVFDVQECQIRGVTYSAPLRVKLRLVIYEREAPEGTVKDIKEQ
EVYMGEIPLMTDNGTFVINGTERVIVSQLHRSPGVFFDSDKGKTHSSGKVLYNARIIPYR
GSWLDFEFDPKDNLFVRIDRRRKLPATIILRALNYTTEQILDLFFEKVIFEIRDNKLQME
LVPERLRGETASFDIEANGKVYVEKGRRITARHIRQLEKDDVKLIEVPVEYIAGKVVAKD
YIDESTGELICAANMELSLDLLAKLSQSGHKRIETLFTNDLDHGPYISETLRVDPTNDRL
SALVEIYRMMRPGEPPTREAAESLFENLFFSEDRYDLSAVGRMKFNRSLLREEIEGSGIL
SKDDIIDVMKKLIDIRNGKGEVDDIDHLGNRRIRSVGEMAENQFRVGLVRVERAVKERLS
LGDLDTLMPQDMINAKPISAAVKEFFGSSQLSQFMDQNNPLSEITHKRRISALGPGGLTR
ERAGFEVRDVHPTHYGRVCPIETPEGPNIGLINSLSVYAQTNEYGFLETPYRKVTDGVVT
DEIHYLSAIEEGNYVIAQANSNLDEEGHFVEDLVTCRSKGESSLFSRDQVDYMDVSTQQV
VSVGASLIPFLEHDDANRALMGANMQRQAVPTLRADKPLVGTGMERAVAVDSGVTAVAKR
GGVVQYVDASRIVIKVNEDEMYPGEAGIDIYNLTKYTRSNQNTCINQMPCVSLGEPVERG
DVLADGPSTDLGELALGQNMRVAFMPWNGYNFEDSILVSERVVQEDRFTTIHIQELACVS
RDTKLGPEEITADIPNVGEAALSKLDESGIVYIGAEVTGGDILVGKVTPKGETQLTPEEK
LLRAIFGEKASDVKDSSLRVPNGVSGTVIDVQVFTRDGVEKDKRALEIEEMQLKQAKKDL
SEELQILEAGLFSRIRAVLVAGGVEAEKLDKLPRDRWLELGLTDEEKQNQLEQLAEQYDE
LKHEFEKKLEAKRRKITQGDDLAPGVLKIVKVYLAVKRRIQPGDKMAGRHGNKGVISKIN
PIEDMPYDENGTPVDIVLNPLGVPSRMNIGQILETHLGMAAKGIGDKINAMLKQQQEVAK
LREFIQRAYDLGADVRQKVDLSTFSDEEVMRLAENLRKGMPIATPVFDGAKEAEIKELLK
LGDLPTSGQIRLYDGRTGEQFERPVTVGYMYMLKLNHLVDDKMHARSTGSYSLVTQQPLG
GKAQFGGQRFGEMEVWALEAYGAAYTLQEMLTVKSDDVNGRTKMYKNIVDGNHQMEPGMP
ESFNVLLKEIRSLGINIELEDE
Sequence of entity 4 (J), FASTA
>6PSV_4 DNA-directed RNA polymerase subunit beta' (chains J)
VKDLLKFLKAQTKTEEFDAIKIALASPDMIRSWSFGEVKKPETINYRTFKPERDGLFCAR
IFGPVKDYECLCGKYKRLKHRGVICEKCGVEVTQTKVRRERMGHIELASPTAHIWFLKSL
PSRIGLLLDMPLRDIERVLYFESYVVIEGGMTNLERQQILTEEQYLDALEEFGDEFDAKM
GAEAIQALLKSMDLEQECEQLREELNETNSETKRKKLTKRIKLLEAFVQSGNKPEWMILT
VLPVLPPDLRPLVPLDGGRFATSDLNDLYRRVINRNNRLKRLLDLAAPDIIVRNEKRMLQ
EAVDALLDNGRRGRAITGSNKRPLKSLADMIKGKQGRFRQNLLGKRVDYSGRSVITVGPY
LRLHQCGLPKKMALELFKPFIYGKLELRGLATTIKAAKKMVEREEAVVWDILDEVIREHP
VLLNRAPTLHRLGIQAFEPVLIEGKAIQLHPLVCAAYNADFDGDQMAVHVPLTLEAQLEA
RALMMSTNNILSPANGEPIIVPSQDVVLGLYYMTRDCVNAKGEGMVLTGPKEAERLYRSG
LASLHARVKVRITEYEKDANGELVAKTSLKDTTVGRAILWMIVPKGLPYSIVNQALGKKA
ISKMLNTCYRILGLKPTVIFADQIMYTGFAYAARSGASVGIDDMVIPEKKHEIISEAEAE
VAEIQEQFQSGLVTAGERYNKVIDIWAAANDRVSKAMMDNLQTETVINRDGQEEKQVSFN
SIYMMADSGARGSAAQIRQLAGMRGLMAKPDGSIIETPITANFREGLNVLQYFISTHGAR
KGLADTALKTANSGYLTRRLVDVAQDLVVTEDDCGTHEGIMMTPVIEGGDVKEPLRDRVL
GRVTAEDVLKPGTADILVPRNTLLHEQWCDLLEENSVDAVKVRSVVSCDTDFGVCAHCYG
RDLARGHIINKGEAIGVIAAQSIGEPGTQLTMRTFHIGGAASRAAAESSIQVKNKGSIKL
SNVKSVVNSSGKLVITSRNTELKLIDEFGRTKESYKVPYGAVLAKGDGEQVAGGETVANW
DPHTMPVITEVSGFVRFTDMIDGQTITRQTDELTGLSSLVVLDSAERTAGGKDLRPALKI
VDAQGNDVLIPGTDMPAQYFLPGKAIVQLEDGVQISSGDTLARIPQESGGTKDITGGLPR
VADLFEARRPKEPAILAEISGIVSFGKETKGKRRLVITPVDGSDPYEEMIPKWRQLNVFE
GERVERGDVISDGPEAPHDILRLRGVHAVTRYIVNEVQDVYRLQGVKINDKHIEVIVRQM
LRKATIVNAGSSDFLEGEQVEYSRVKIANRELEANGKVGATYSRDLLGITKASLATESFI
SAASFQETTRVLTEAAVAGKRDELRGLKENVIVGRLIPAGTGYAYHQDRMRRRAAGEAPA
APQVTAEDASASLAELLNAGLGGSDNELELEVLFQGPSSGHHHHHHHHHH
Sequence of entity 5 (K), FASTA
>6PSV_5 DNA-directed RNA polymerase subunit omega (chains K)
MARVTVQDAVEKIGNRFDLVLVAARRARQMQVGGKDPLVPEENDKTTVIALREIEEGLIN
NQILDVRERQEQQEQEAAELQAVTAIAEGRR
Sequence of entity 6 (L), FASTA
>6PSV_6 RNA polymerase sigma factor RpoD (chains L)
SEFMEQNPQSQLKLLVTRGKEQGYLTYAEVNDHLPEDIVDSDQIEDIIQMINDMGIQVME
EAPDADDLMLAENTADEDAAEAAAQVLSSVESEIGRTTDPVRMYMREMGTVELLTREGEI
DIAKRIEDGINQVQCSVAEYPEAITYLLEQYDRVEAEEARLSDLITGFVDPNAEEDLAPT
ATHVGSELSQEDLDDDEDEDEEDGDDDSADDDNSIDPELAREKFAELRAQYVVTRDTIKA
KGRSHATAQEEILKLSEVFKQFRLVPKQFDYLVNSMRVMMDRVRTQERLIMKLCVEQCKM
PKKNFITLFTGNETSDTWFNAAIAMNKPWSEKLHDVSEEVHRALQKLQQIEEETGLTIEQ
VKDINRRMSIGEAKARRAKKEMVEANLRLVISIAKKYTNRGLQFLDLIQEGNIGLMKAVD
KFEYRRGYKFSTYATWWIRQAITRSIADQARTIRIPVHMIETINKLNRISRQMLQEMGRE
PTPEELAERMLMPEDKIRKVLKIAKEPISMETPIGDDEDSHLGDFIEDTTLELPLDSATT
ESLRAATHDVLAGLTAREAKVLRMRFGIDMNTDYTLEEVGKQFDVTRERIRQIEAKALRK
LRHPSRSEVLRSFLDD
Sequence of entity 7 (O), FASTA
>6PSV_7 DNA (85-MER) (chains O)
GGCGGCGCTTATTTGCACAAATCCATTGACAAAAGAAGGCTAAAAGGGCATATTCCTCGG
CCTTTGAATTGTCCATATAGAACGC
Sequence of entity 8 (P), FASTA
>6PSV_8 DNA (85-MER) (chains P)
GCGTTCTATATGGACAATTCAAAGGCCGAGGAATATGCCCTTTTAGCCTTCTTTTGTCAA
TGGATTTGTGCAAATAAGCGCCGCC

Ligands and cofactors

IDNameFormulaCopies
ZNZinc ionZn3
1N7ChapsoC32 H59 N2 O8 S4
MGMagnesium ionMg1

Primary citation

Stepwise Promoter Melting by Bacterial RNA Polymerase. Chen, J., Chiu, C., Gopalkrishnan, S. et al. Mol Cell (2020) 78:275. DOI 10.1016/j.molcel.2020.02.017 · PubMed

Other PDB entries of the same protein (UniProt P41065 (AlphaFold model), which also has an AlphaFold model), best resolution first:

Browse structure collections

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