9R3D: Human pre-Bact-OTS complex
Cryo-EM structure of the human pre-Bact-OTS complex (whole map). Determined by electron microscopy at 3.12 Å resolution. Released 13 May 2026.
- Method
- Electron microscopy
- Resolution
- 3.12 Å
- Organism
- Homo sapiens
- Chains
- 17
- Atoms
- 41,862
- Mol. weight
- 1255.13 kDa
- Ligands
- IHP
- Released
- 13 May 2026
Explore 9R3D in 3D
Show helices and sheets
RCSB PDB
PDBe
Secondary structure: helices and β-sheets
9R3D contains 257 α-helices and 158 β-strands across 14 chains. Residue ranges use author residue numbering, as in the PDB file, from PDBe. To see them in 3D, choose the Cartoon representation with Secondary structure coloring: helices, sheets and coils get different colors.
Chain A: 110 helices, 64 β-strands
| Element | Residues | Length | Sheet |
|---|
| β-strand | 60 | 1 | 26 |
| α-helix | 61-62 | 2 | |
| α-helix | 63-73 | 11 | |
| α-helix | 85-90 | 6 | |
| α-helix | 91-93 | 3 | |
| α-helix | 95-104 | 10 | |
| α-helix | 106-108 | 3 | |
| β-strand | 115-121 | 7 | 26 |
| β-strand | 127-129 | 3 | 26 |
| β-strand | 134-135 | 2 | 27 |
| α-helix | 137-157 | 21 | |
| α-helix | 163-165 | 3 | |
| α-helix | 173-176 | 4 | |
| α-helix | 178 | 1 | |
| α-helix | 179-183 | 5 | |
| α-helix | 186-190 | 5 | |
| β-strand | 220 | 1 | 28 |
| β-strand | 227-229 | 3 | 28 |
| α-helix | 232-241 | 10 | |
| α-helix | 252-255 | 4 | |
| α-helix | 260-268 | 9 | |
| α-helix | 287-290 | 4 | |
| α-helix | 299-301 | 3 | |
| α-helix | 310-315 | 6 | |
| α-helix | 317-320 | 4 | |
| α-helix | 329-331 | 3 | |
| α-helix | 357-359 | 3 | |
| α-helix | 396-404 | 9 | |
| α-helix | 409-412 | 4 | |
| β-strand | 415-417 | 3 | 28 |
| α-helix | 418 | 1 | |
| β-strand | 419-420 | 2 | 27 |
| α-helix | 421-423 | 3 | |
| α-helix | 428-432 | 5 | |
| α-helix | 434-436 | 3 | |
| α-helix | 441-459 | 19 | |
| α-helix | 463-466 | 4 | |
| α-helix | 472-478 | 7 | |
| β-strand | 482-488 | 7 | 26 |
| α-helix | 489-510 | 22 | |
| β-strand | 516-519 | 4 | 29 |
| β-strand | 523-526 | 4 | 29 |
| α-helix | 532-537 | 6 | |
| α-helix | 542-566 | 25 | |
| α-helix | 571-583 | 13 | |
| α-helix | 585-588 | 4 | |
| α-helix | 591-593 | 3 | |
| α-helix | 598-616 | 19 | |
| α-helix | 631-660 | 30 | |
| α-helix | 678-696 | 19 | |
| α-helix | 698-701 | 4 | |
| α-helix | 706-722 | 17 | |
| α-helix | 733-763 | 31 | |
| α-helix | 769-796 | 28 | |
| α-helix | 803-819 | 17 | |
| α-helix | 822-829 | 8 | |
| α-helix | 835-850 | 16 | |
| α-helix | 858-872 | 15 | |
| α-helix | 874-886 | 13 | |
| α-helix | 892-893 | 2 | |
| β-strand | 894 | 1 | 30 |
| α-helix | 895 | 1 | |
| β-strand | 897-900 | 4 | 31 |
| β-strand | 905-908 | 4 | 31 |
| α-helix | 913-932 | 20 | |
| β-strand | 940 | 1 | 32 |
| α-helix | 948-961 | 14 | |
| β-strand | 973-980 | 8 | 32 |
| β-strand | 981 | 1 | 33 |
| α-helix | 990-1000 | 11 | |
| α-helix | 1003-1013 | 11 | |
| β-strand | 1016-1017 | 2 | 30 |
| β-strand | 1024-1025 | 2 | 30 |
| α-helix | 1037-1053 | 17 | |
| α-helix | 1055-1062 | 8 | |
| α-helix | 1077-1081 | 5 | |
| β-strand | 1085-1090 | 6 | 32 |
| β-strand | 1094-1101 | 8 | 32 |
| α-helix | 1103-1116 | 14 | |
| α-helix | 1144-1159 | 16 | |
| β-strand | 1168 | 1 | 33 |
| α-helix | 1170-1172 | 3 | |
| β-strand | 1174-1178 | 5 | 32 |
| β-strand | 1185-1189 | 5 | 34 |
| β-strand | 1192-1198 | 7 | 34 |
| β-strand | 1213-1217 | 5 | 34 |
| β-strand | 1224-1232 | 9 | 34 |
| α-helix | 1234-1248 | 15 | |
| α-helix | 1256-1271 | 16 | |
| α-helix | 1275-1278 | 4 | |
| α-helix | 1282-1303 | 22 | |
| α-helix | 1307-1308 | 2 | |
| α-helix | 1313-1317 | 5 | |
| α-helix | 1320-1322 | 3 | |
| α-helix | 1327-1329 | 3 | |
| β-strand | 1334 | 1 | 35 |
| α-helix | 1337-1339 | 3 | |
| β-strand | 1341-1345 | 5 | 36 |
| β-strand | 1350-1354 | 5 | 36 |
| β-strand | 1364 | 1 | 35 |
| α-helix | 1365-1367 | 3 | |
| α-helix | 1368-1371 | 4 | |
| α-helix | 1375-1399 | 25 | |
| α-helix | 1406-1411 | 6 | |
| α-helix | 1419-1421 | 3 | |
| α-helix | 1427-1430 | 4 | |
| α-helix | 1436-1442 | 7 | |
| α-helix | 1443-1445 | 3 | |
| α-helix | 1458-1461 | 4 | |
| α-helix | 1468-1478 | 11 | |
| α-helix | 1480-1487 | 8 | |
| α-helix | 1491-1493 | 3 | |
| β-strand | 1501-1503 | 3 | 37 |
| α-helix | 1531-1537 | 7 | |
| α-helix | 1539-1542 | 4 | |
| β-strand | 1553-1554 | 2 | 38 |
| β-strand | 1561-1562 | 2 | 38 |
| α-helix | 1567-1574 | 8 | |
| α-helix | 1581-1598 | 18 | |
| β-strand | 1608-1611 | 4 | 39 |
| α-helix | 1616-1620 | 5 | |
| β-strand | 1629-1637 | 9 | 39 |
| α-helix | 1639-1642 | 4 | |
| β-strand | 1643-1644 | 2 | 39 |
| α-helix | 1655-1656 | 2 | |
| β-strand | 1657-1667 | 11 | 39 |
| α-helix | 1676-1687 | 12 | |
| β-strand | 1700-1706 | 7 | 39 |
| β-strand | 1711-1716 | 6 | 39 |
| α-helix | 1722-1735 | 14 | |
| α-helix | 1738-1751 | 14 | |
| β-strand | 1753-1755 | 3 | 37 |
| α-helix | 1796-1798 | 3 | |
| β-strand | 1811 | 1 | 40 |
| β-strand | 1817 | 1 | 40 |
| α-helix | 1832-1850 | 19 | |
| α-helix | 1869-1878 | 10 | |
| α-helix | 1897-1907 | 11 | |
| α-helix | 1929-1945 | 17 | |
| α-helix | 1947-1952 | 6 | |
| α-helix | 1973-1994 | 22 | |
| α-helix | 2004-2009 | 6 | |
| β-strand | 2040-2047 | 8 | 23 |
| β-strand | 2053-2058 | 6 | 23 |
| β-strand | 2068-2070 | 3 | 39 |
| α-helix | 2074-2086 | 13 | |
| α-helix | 2095-2098 | 4 | |
| α-helix | 2101-2103 | 3 | |
| β-strand | 2104-2105 | 2 | 41 |
| β-strand | 2107 | 1 | 42 |
| α-helix | 2108-2116 | 9 | |
| β-strand | 2125-2128 | 4 | 43 |
| β-strand | 2131 | 1 | 41 |
| α-helix | 2132-2133 | 2 | |
| β-strand | 2140-2141 | 2 | 41 |
| β-strand | 2144-2146 | 3 | 43 |
| β-strand | 2151-2152 | 2 | 44 |
| β-strand | 2157-2158 | 2 | 44 |
| α-helix | 2167-2171 | 5 | |
| β-strand | 2177-2179 | 3 | 43 |
| β-strand | 2180-2181 | 2 | 45 |
| α-helix | 2190-2202 | 13 | |
| β-strand | 2216-2217 | 2 | 45 |
| β-strand | 2228 | 1 | 46 |
| α-helix | 2235-2239 | 5 | |
| α-helix | 2253-2255 | 3 | |
| β-strand | 2258 | 1 | 46 |
| β-strand | 2261 | 1 | 41 |
| β-strand | 2265 | 1 | 42 |
| β-strand | 2270-2272 | 3 | 43 |
| β-strand | 2273-2274 | 2 | 47 |
| α-helix | 2282-2284 | 3 | |
| β-strand | 2296-2297 | 2 | 47 |
| α-helix | 2313-2319 | 7 | |
Chain B: 39 helices, 19 β-strands
| Element | Residues | Length | Sheet |
|---|
| β-strand | 445 | 1 | 1 |
| α-helix | 469-472 | 4 | |
| α-helix | 483-494 | 12 | |
| β-strand | 502 | 1 | 2 |
| α-helix | 510-524 | 15 | |
| α-helix | 526-528 | 3 | |
| α-helix | 545-561 | 17 | |
| α-helix | 577-581 | 5 | |
| α-helix | 590-599 | 10 | |
| α-helix | 601-604 | 4 | |
| α-helix | 624-634 | 11 | |
| α-helix | 638-641 | 4 | |
| β-strand | 652 | 1 | 2 |
| α-helix | 660-665 | 6 | |
| β-strand | 685 | 1 | 3 |
| β-strand | 688-690 | 3 | 1 |
| α-helix | 697-714 | 18 | |
| α-helix | 730-747 | 18 | |
| α-helix | 755-757 | 3 | |
| α-helix | 763-767 | 5 | |
| α-helix | 792-802 | 11 | |
| α-helix | 816-819 | 4 | |
| β-strand | 828-829 | 2 | 1 |
| α-helix | 847-854 | 8 | |
| β-strand | 865 | 1 | 3 |
| β-strand | 868-870 | 3 | 1 |
| α-helix | 875-879 | 5 | |
| α-helix | 886-890 | 5 | |
| α-helix | 903-907 | 5 | |
| α-helix | 908-910 | 3 | |
| α-helix | 914-935 | 22 | |
| α-helix | 939-943 | 5 | |
| α-helix | 950-953 | 4 | |
| α-helix | 955-981 | 27 | |
| α-helix | 983-990 | 8 | |
| α-helix | 995-1004 | 10 | |
| α-helix | 1011-1020 | 10 | |
| α-helix | 1034-1042 | 9 | |
| α-helix | 1055-1068 | 14 | |
| α-helix | 1075-1101 | 27 | |
| α-helix | 1105-1120 | 16 | |
| α-helix | 1128-1131 | 4 | |
| α-helix | 1137-1145 | 9 | |
| α-helix | 1158-1165 | 8 | |
| α-helix | 1168-1180 | 13 | |
| β-strand | 1186 | 1 | 4 |
| β-strand | 1189-1191 | 3 | 5 |
| β-strand | 1199-1203 | 5 | 5 |
| β-strand | 1204 | 1 | 4 |
| α-helix | 1212-1215 | 4 | |
| β-strand | 1219-1221 | 3 | 6 |
| β-strand | 1222-1227 | 6 | 7 |
| β-strand | 1233-1236 | 4 | 7 |
| β-strand | 1239-1241 | 3 | 6 |
| α-helix | 1243-1246 | 4 | |
| β-strand | 1251-1254 | 4 | 5 |
| β-strand | 1266-1271 | 6 | 7 |
| β-strand | 1280-1283 | 4 | 7 |
Chain C: 40 helices, 39 β-strands
| Element | Residues | Length | Sheet |
|---|
| α-helix | 116-124 | 9 | |
| β-strand | 129-133 | 5 | 8 |
| β-strand | 136 | 1 | 8 |
| α-helix | 142-153 | 12 | |
| α-helix | 172-177 | 6 | |
| β-strand | 184-191 | 8 | 8 |
| β-strand | 197-204 | 8 | 8 |
| α-helix | 212-221 | 10 | |
| β-strand | 224-230 | 7 | 8 |
| α-helix | 237-248 | 12 | |
| β-strand | 252-258 | 7 | 8 |
| α-helix | 261 | 1 | |
| α-helix | 262-266 | 5 | |
| α-helix | 271-292 | 22 | |
| β-strand | 307-311 | 5 | 8 |
| β-strand | 316-318 | 3 | 8 |
| α-helix | 322-330 | 9 | |
| α-helix | 336-341 | 6 | |
| β-strand | 348-350 | 3 | 9 |
| β-strand | 355-357 | 3 | 9 |
| α-helix | 365-366 | 2 | |
| β-strand | 367 | 1 | 9 |
| α-helix | 368-369 | 2 | |
| α-helix | 370-374 | 5 | |
| α-helix | 375-386 | 12 | |
| α-helix | 392-399 | 8 | |
| α-helix | 405-408 | 4 | |
| α-helix | 412-424 | 13 | |
| α-helix | 428-437 | 10 | |
| α-helix | 441-452 | 12 | |
| β-strand | 453 | 1 | 10 |
| α-helix | 460-466 | 7 | |
| β-strand | 475-483 | 9 | 11 |
| β-strand | 490-495 | 6 | 11 |
| β-strand | 501-502 | 2 | 12 |
| β-strand | 506-510 | 5 | 11 |
| β-strand | 522-526 | 5 | 11 |
| β-strand | 530-534 | 5 | 11 |
| β-strand | 537-541 | 5 | 11 |
| β-strand | 543-544 | 2 | 12 |
| β-strand | 549-552 | 4 | 11 |
| β-strand | 563-566 | 4 | 11 |
| β-strand | 577 | 1 | 10 |
| α-helix | 578-580 | 3 | |
| β-strand | 588-594 | 7 | 13 |
| α-helix | 597-599 | 3 | |
| α-helix | 600-613 | 14 | |
| β-strand | 618-621 | 4 | 13 |
| β-strand | 627-631 | 5 | 13 |
| α-helix | 634-642 | 9 | |
| α-helix | 643-649 | 7 | |
| β-strand | 653-655 | 3 | 13 |
| α-helix | 656-658 | 3 | |
| β-strand | 659-660 | 2 | 13 |
| β-strand | 663-666 | 4 | 14 |
| β-strand | 674-677 | 4 | 15 |
| β-strand | 684-691 | 8 | 15 |
| α-helix | 696-701 | 6 | |
| α-helix | 717-720 | 4 | |
| α-helix | 726-729 | 4 | |
| β-strand | 735 | 1 | 15 |
| β-strand | 743-747 | 5 | 15 |
| α-helix | 757-776 | 20 | |
| β-strand | 784 | 1 | 14 |
| β-strand | 787-794 | 8 | 15 |
| α-helix | 801-803 | 3 | |
| α-helix | 805-822 | 18 | |
| β-strand | 825-839 | 15 | 14 |
| α-helix | 840-842 | 3 | |
| α-helix | 843-852 | 10 | |
| β-strand | 856-862 | 7 | 14 |
| α-helix | 863 | 1 | |
| β-strand | 868-876 | 9 | 14 |
| α-helix | 883-890 | 8 | |
| β-strand | 896-906 | 11 | 14 |
| α-helix | 907 | 1 | |
| α-helix | 923-925 | 3 | |
| α-helix | 926-928 | 3 | |
| α-helix | 929-940 | 12 | |
Chain I: 14 helices, 9 β-strands
| Element | Residues | Length | Sheet |
|---|
| α-helix | 5-6 | 2 | |
| β-strand | 12 | 1 | 16 |
| β-strand | 15 | 1 | 16 |
| α-helix | 17-20 | 4 | |
| α-helix | 23-30 | 8 | |
| α-helix | 33-37 | 5 | |
| α-helix | 44-51 | 8 | |
| β-strand | 57-58 | 2 | 17 |
| β-strand | 60-61 | 2 | 18 |
| β-strand | 66-67 | 2 | 18 |
| α-helix | 69-80 | 12 | |
| α-helix | 84-92 | 9 | |
| α-helix | 97-109 | 13 | |
| α-helix | 113-119 | 7 | |
| α-helix | 121-125 | 5 | |
| β-strand | 129-133 | 5 | 17 |
| β-strand | 139-143 | 5 | 17 |
| α-helix | 144-153 | 10 | |
| β-strand | 156-157 | 2 | 19 |
| β-strand | 160-161 | 2 | 19 |
| α-helix | 162-164 | 3 | |
| α-helix | 168-173 | 6 | |
| α-helix | 178-180 | 3 | |
Chain K: 7 helices, 0 β-strands
| Element | Residues | Length | Sheet |
|---|
| α-helix | 272-317 | 46 | |
| α-helix | 335-336 | 2 | |
| α-helix | 339-341 | 3 | |
| α-helix | 354-356 | 3 | |
| α-helix | 365-368 | 4 | |
| α-helix | 376-378 | 3 | |
| α-helix | 394-397 | 4 | |
Chain P: 13 helices, 7 β-strands
| Element | Residues | Length | Sheet |
|---|
| α-helix | 20-22 | 3 | |
| α-helix | 25-28 | 4 | |
| β-strand | 33-39 | 7 | 25 |
| α-helix | 53 | 1 | |
| β-strand | 54-57 | 4 | 25 |
| β-strand | 66-67 | 2 | 25 |
| α-helix | 72-77 | 6 | |
| β-strand | 80 | 1 | 48 |
| β-strand | 87 | 1 | 48 |
| α-helix | 94-98 | 5 | |
| α-helix | 146-149 | 4 | |
| α-helix | 157-160 | 4 | |
| α-helix | 162-163 | 2 | |
| β-strand | 164 | 1 | 49 |
| α-helix | 165 | 1 | |
| α-helix | 166-169 | 4 | |
| α-helix | 176-178 | 3 | |
| β-strand | 182 | 1 | 49 |
| α-helix | 199-203 | 5 | |
| α-helix | 208-219 | 12 | |
Chain P7: 1 helix, 1 β-strand
| Element | Residues | Length | Sheet |
|---|
| β-strand | 115-117 | 3 | 25 |
| α-helix | 125-130 | 6 | |
Chain q: 4 helices, 7 β-strands
| Element | Residues | Length | Sheet |
|---|
| β-strand | 2-7 | 6 | 20 |
| β-strand | 13-18 | 6 | 20 |
| β-strand | 23 | 1 | 21 |
| α-helix | 24-35 | 12 | |
| α-helix | 39-41 | 3 | |
| β-strand | 42-46 | 5 | 20 |
| β-strand | 49-50 | 2 | 20 |
| α-helix | 51 | 1 | |
| β-strand | 56 | 1 | 21 |
| α-helix | 57-60 | 4 | |
| β-strand | 67-72 | 6 | 20 |
6 more chain groups are not listed. Open the entry in the viewer and use the sequence panel to see them.
Molecules and chains
| Molecule | Chains | Type | Length | Organism | UniProt |
|---|
| U5 snRNA | 5 | RNA | 77 | Homo sapiens | |
| U6 snRNA | 6 | RNA | 85 | Homo sapiens | |
| U5 small nuclear ribonucleoprotein 200 kDa helicase | B | protein | 2136 | Homo sapiens | O75643 (AlphaFold model) |
| 116 kDa U5 small nuclear ribonucleoprotein component | C | protein | 972 | Homo sapiens | Q15029 (AlphaFold model) |
| Pre-mRNA-splicing factor 38A | I | protein | 312 | Homo sapiens | Q8NAV1 (AlphaFold model) |
| Microfibrillar-associated protein 1 | K | protein | 439 | Homo sapiens | P55081 (AlphaFold model) |
| Protein BUD31 homolog | Q | protein | 144 | Homo sapiens | P41223 |
| Transcription elongation regulator 1 | T | protein | 1098 | Homo sapiens | O14776 |
| WW domain-binding protein 11 | X | protein | 641 | Homo sapiens | Q9Y2W2 |
| pre-mRNA | Z | RNA | 29 | Homo sapiens | |
| Ubiquitin-like protein 5 | q | protein | 73 | Homo sapiens | Q9BZL1 |
| Zinc finger matrin-type protein 2 | r | protein | 199 | Homo sapiens | Q96NC0 |
5 more molecules are not listed.
Sequence of entity 1 (5), FASTA
>9R3D_1 U5 snRNA (chains 5)
ACUCUGGUUUCUCUUCAGAUCGCAUAAAUCUUUCGCCUUUUACUAAAGAUUUCCGUGGAG
AGGAACAACUCUGAGUC
Sequence of entity 2 (6), FASTA
>9R3D_2 U6 snRNA (chains 6)
GUGCUCGCUUCGGCAGCACAUAUACUAAAAUUGGAACGAUACAGAGAAGAUUAGCAUGGC
CCCUGCGCAAGGAUGACACGCAAAA
Sequence of entity 3 (B), FASTA
>9R3D_3 U5 small nuclear ribonucleoprotein 200 kDa helicase (chains B)
MADVTARSLQYEYKANSNLVLQADRSLIDRTRRDEPTGEVLSLVGKLEGTRMGDKAQRTK
PQMQEERRAKRRKRDEDRHDINKMKGYTLLSEGIDEMVGIIYKPKTKETRETYEVLLSFI
QAALGDQPRDILCGAADEVLAVLKNEKLRDKERRKEIDLLLGQTDDTRYHVLVNLGKKIT
DYGGDKEIQNMDDNIDETYGVNVQFESDEEEGDEDVYGEVREEASDDDMEGDEAVVRCTL
SANLVASGELMSSKKKDLHPRDIDAFWLQRQLSRFYDDAIVSQKKADEVLEILKTASDDR
ECENQLVLLLGFNTFDFIKVLRQHRMMILYCTLLASAQSEAEKERIMGKMEADPELSKFL
YQLHETEKEDLIREERSRRERVRQSRMDTDLETMDLDQGGEALAPRQVLDLEDLVFTQGS
HFMANKRCQLPDGSFRRQRKGYEEVHVPALKPKPFGSEEQLLPVEKLPKYAQAGFEGFKT
LNRIQSKLYRAALETDENLLLCAPTGAGKTNVALMCMLREIGKHINMDGTINVDDFKIIY
IAPMRSLVQEMVGSFGKRLATYGITVAELTGDHQLCKEEISATQIIVCTPEKWDIITRKG
GERTYTQLVRLIILDEIHLLHDDRGPVLEALVARAIRNIEMTQEDVRLIGLSATLPNYED
VATFLRVDPAKGLFYFDNSFRPVPLEQTYVGITEKKAIKRFQIMNEIVYEKIMEHAGKNQ
VLVFVHSRKETGKTARAIRDMCLEKDTLGLFLREGSASTEVLRTEAEQCKNLELKDLLPY
GFAIHHAGMTRVDRTLVEDLFADKHIQVLVSTATLAWGVNLPAHTVIIKGTQVYSPEKGR
WTELGALDILQMLGRAGRPQYDTKGEGILITSHGELQYYLSLLNQQLPIESQMVSKLPDM
LNAEIVLGNVQNAKDAVNWLGYAYLYIRMLRSPTLYGISHDDLKGDPLLDQRRLDLVHTA
ALMLDKNNLVKYDKKTGNFQVTELGRIASHYYITNDTVQTYNQLLKPTLSEIELFRVFSL
SSEFKNITVREEEKLELQKLLERVPIPVKESIEEPSAKINVLLQAFISQLKLEGFALMAD
MVYVTQSAGRLMRAIFEIVLNRGWAQLTDKTLNLCKMIDKRMWQSMCPLRQFRKLPEEVV
KKIEKKNFPFERLYDLNHNEIGELIRMPKMGKTIHKYVHLFPKLELSVHLQPITRSTLKV
ELTITPDFQWDEKVHGSSEAFWILVEDVDSEVILHHEYFLLKAKYAQDEHLITFFVPVFE
PLPPQYFIRVVSDRWLSCETQLPVSFRHLILPEKYPPPTELLDLQPLPVSALRNSAFESL
YQDKFPFFNPIQTQVFNTVYNSDDNVFVGAPTGSGKTICAEFAILRMLLQSSEGRCVYIT
PMEALAEQVYMDWYEKFQDRLNKKVVLLTGETSTDLKLLGKGNIIISTPEKWDILSRRWK
QRKNVQNINLFVVDEVHLIGGENGPVLEVICSRMRYISSQIERPIRIVALSSSLSNAKDV
AHWLGCSATSTFNFHPNVRPVPLELHIQGFNISHTQTRLLSMAKPVYHAITKHSPKKPVI
VFVPSRKQTRLTAIDILTTCAADIQRQRFLHCTEKDLIPYLEKLSDSTLKETLLNGVGYL
HEGLSPMERRLVEQLFSSGAIQVVVASRSLCWGMNVAAHLVIIMDTQYYNGKIHAYVDYP
IYDVLQMVGHANRPLQDDEGRCVIMCQGSKKDFFKKFLYEPLPVESHLDHCMHDHFNAEI
VTKTIENKQDAVDYLTWTFLYRRMTQNPNYYNLQGISHRHLSDHLSELVEQTLSDLEQSK
CISIEDEMDVAPLNLGMIAAYYYINYTTIELFSMSLNAKTKVRGLIEIISNAAEYENIPI
RHHEDNLLRQLAQKVPHKLNNPKFNDPHVKTNLLLQAHLSRMQLSAELQSDTEEILSKAI
RLIQACVDVLSSNGWLSPALAAMELAQMVTQAMWSKDSYLKQLPHFTSEHIKRCTDKGVE
SVFDIMEMEDEERNALLQLTDSQIADVARFCNRYPNIELSYEVVDKDSIRSGGPVVVLVQ
LEREEEVTGPVIAPLFPQKREEGWWVVIGDAKSNSLISIKRLTLQQKAKVKLDFVAPATG
AHNYTLYFMSDAYMGCDQEYKFSVDVKEAETDSDSD
Sequence of entity 4 (C), FASTA
>9R3D_4 116 kDa U5 small nuclear ribonucleoprotein component (chains C)
MDTDLYDEFGNYIGPELDSDEDDDELGRETKDLDEMDDDDDDDDVGDHDDDHPGMEVVLH
EDKKYYPTAEEVYGPEVETIVQEEDTQPLTEPIIKPVKTKKFTLMEQTLPVTVYEMDFLA
DLMDNSELIRNVTLCGHLHHGKTCFVDCLIEQTHPEIRKRYDQDLCYTDILFTEQERGVG
IKSTPVTVVLPDTKGKSYLFNIMDTPGHVNFSDEVTAGLRISDGVVLFIDAAEGVMLNTE
RLIKHAVQERLAVTVCINKIDRLILELKLPPTDAYYKLRHIVDEVNGLISMYSTDENLIL
SPLLGNVCFSSSQYSICFTLGSFAKIYADTFGDINYQEFAKRLWGDIYFNPKTRKFTKKA
PTSSSQRSFVEFILEPLYKILAQVVGDVDTSLPRTLDELGIHLTKEELKLNIRPLLRLVC
KKFFGEFTGFVDMCVQHIPSPKVGAKPKIEHTYTGGVDSDLGEAMSDCDPDGPLMCHTTK
MYSTDDGVQFHAFGRVLSGTIHAGQPVKVLGENYTLEDEEDSQICTVGRLWISVARYHIE
VNRVPAGNWVLIEGVDQPIVKTATITEPRGNEEAQIFRPLKFNTTSVIKIAVEPVNPSEL
PKMLDGLRKVNKSYPSLTTKVEESGEHVILGTGELYLDCVMHDLRKMYSEIDIKVADPVV
TFCETVVETSSLKCFAETPNKKNKITMIAEPLEKGLAEDIENEVVQITWNRKKLGEFFQT
KYDWDLLAARSIWAFGPDATGPNILVDDTLPSEVDKALLGSVKDSIVQGFQWGTREGPLC
DELIRNVKFKILDAVVAQEPLHRGGGQIIPTARRVVYSAFLMATPRLMEPYYFVEVQAPA
DCVSAVYTVLARRRGHVTQDAPIPGSPLYTIKAFIPAIDSFGFETDLRTHTQGQAFSLSV
FHHWQIVPGDPLDKSIVIRPLEPQPAPHLAREFMIKTRRRKGLSEDVSISKFFDDPMLLE
LAKQDVVLNYPM
Sequence of entity 5 (I), FASTA
>9R3D_5 Pre-mRNA-splicing factor 38A (chains I)
MANRTVKDAHSIHGTNPQYLVEKIIRTRIYESKYWKEECFGLTAELVVDKAMELRFVGGV
YGGNIKPTPFLCLTLKMLQIQPEKDIIVEFIKNEDFKYVRMLGALYMRLTGTAIDCYKYL
EPLYNDYRKIKSQNRNGEFELMHVDEFIDELLHSERVCDIILPRLQKRYVLEEAEQLEPR
VSALEEDMDDVESSEEEEEEDEKLERVPSPDHRRRSYRDLDKPRRSPTLRYRRSRSRSPR
RRSRSPKRRSPSPRRERHRSKSPRRHRSRSRDRRHRSRSKSPGHHRSHRHRSHSKSPERS
KKSHKKSRRGNE
Sequence of entity 6 (K), FASTA
>9R3D_6 Microfibrillar-associated protein 1 (chains K)
MSVPSALMKQPPIQSTAGAVPVRNEKGEISMEKVKVKRYVSGKRPDYAPMESSDEEDEEF
QFIKKAKEQEAEPEEQEEDSSSDPRLRRLQNRISEDVEERLARHRKIVEPEVVGESDSEV
EGDAWRMEREDSSEEEEEEIDDEEIERRRGMMRQRAQERKNEEMEVMEVEDEGRSGEESE
SESEYEEYTDSEDEMEPRLKPVFIRKKDRVTVQEREAEALKQKELEQEAKRMAEERRKYT
LKIVEEETKKELEENKRSLAALDALNTDDENDEEEYEAWKVRELKRIKRDREDREALEKE
KAEIERMRNLTEEERRAELRANGKVITNKAVKGKYKFLQKYYHRGAFFMDEDEEVYKRDF
SAPTLEDHFNKTILPKVMQVKNFGRSGRTKYTHLVDQDTTSFDSAWGQESAQNTKFFKQK
AAGVRDVFERPSAKKRKTT
Sequence of entity 7 (Q), FASTA
>9R3D_7 Protein BUD31 homolog (chains Q)
MPKVKRSRKAPPDGWELIEPTLDELDQKMREAETEPHEGKRKVESLWPIFRIHHQKTRYI
FDLFYKRKAISRELYEYCIKEGYADKNLIAKWKKQGYENLCCLRCIQTRDTNFGTNCICR
VPKSKLEVGRIIECTHCGCRGCSG
Sequence of entity 8 (T), FASTA
>9R3D_8 Transcription elongation regulator 1 (chains T)
MAERGGDGGESERFNPGELRMAQQQALRFRGPAPPPNAVMRGPPPLMRPPPPFGMMRGPP
PPPRPPFGRPPFDPNMPPMPPPGGIPPPMGPPHLQRPPFMPPPMSSMPPPPGMMFPPGMP
PVTAPGTPALPPTEEIWVENKTPDGKVYYYNARTRESAWTKPDGVKVIQQSELTPMLAAQ
AQVQAQAQAQAQAQAQAQAQAQAQAQAQAQAQAQAQAQAQAQAQAQAQAQAQAQAQAQAQ
AQVQAQVQAQVQAQAVGASTPTTSSPAPAVSTSTSSSTPSSTTSTTTTATSVAQTVSTPT
TQDQTPSSAVSVATPTVSVSTPAPTATPVQTVPQPHPQTLPPAVPHSVPQPTTAIPAFPP
VMVPPFRVPLPGMPIPLPGVAMMQIVSCPYVKTVATTKTGVLPGMAPPIVPMIHPQVAIA
ASPATLAGATAVSEWTEYKTADGKTYYYNNRTLESTWEKPQELKEKEKLEEKIKEPIKEP
SEEPLPMETEEEDPKEEPIKEIKEEPKEEEMTEEEKAAQKAKPVATAPIPGTPWCVVWTG
DERVFFYNPTTRLSMWDRPDDLIGRADVDKIIQEPPHKKGMEELKKLRHPTPTMLSIQKW
QFSMSAIKEEQELMEEINEDEPVKAKKRKRDDNKDIDSEKEAAMEAEIKAARERAIVPLE
ARMKQFKDMLLERGVSAFSTWEKELHKIVFDPRYLLLNPKERKQVFDQYVKTRAEEERRE
KKNKIMQAKEDFKKMMEEAKFNPRATFSEFAAKHAKDSRFKAIEKMKDREALFNEFVAAA
RKKEKEDSKTRGEKIKSDFFELLSNHHLDSQSRWSKVKDKVESDPRYKAVDSSSMREDLF
KQYIEKIAKNLDSEKEKELERQARIEASLREREREVQKARSEQTKEIDREREQHKREEAI
QNFKALLSDMVRSSDVSWSDTRRTLRKDHRWESGSLLEREEKEKLFNEHIEALTKKKREH
FRQLLDETSAITLTSTWKEVKKIIKEDPRCIKFSSSDRKKQREFEEYIRDKYITAKADFR
TLLKETKFITYRSKKLIQESDQHLKDVEKILQNDKRYLVLDCVPEERRKLIVAYVDDLDR
RGPPPPPTASEPTRRSTK
Sequence of entity 9 (X), FASTA
>9R3D_9 WW domain-binding protein 11 (chains X)
MGRRSTSSTKSGKFMNPTDQARKEARKRELKKNKKQRMMVRAAVLKMKDPKQIIRDMEKL
DEMEFNPVQQPQLNEKVLKDKRKKLRETFERILRLYEKENPDIYKELRKLEVEYEQKRAQ
LSQYFDAVKNAQHVEVESIPLPDMPHAPSNILIQDIPLPGAQPPSILKKTSAYGPPTRAV
SILPLLGHGVPRLPPGRKPPGPPPGPPPPQVVQMYGRKVGFALDLPPRRRDEDMLYSPEL
AQRGHDDDVSSTSEDDGYPEDMDQDKHDDSTDDSDTDKSDGESDGDEFVHRDNGERDNNE
EKKSGLSVRFADMPGKSRKKKKNMKELTPLQAMMLRMAGQEIPEEGREVEEFSEDDDEDD
SDDSEAEKQSQKQHKEESHSDGTSTASSQQQAPPQSVPPSQIQAPPMPGPPPLGPPPAPP
LRPPGPPTGLPPGPPPGAPPFLRPPGMPGLRGPLPRLLPPGPPPGRPPGPPPGPPPGLPP
GPPPRGPPPRLPPPAPPGIPPPRPGMMRPPLVPPLGPAPPGLFPPAPLPNPGVLSAPPNL
IQRPKADDTSAATIEKKATATISAKPQITNPKAEITRFVPTALRVRRENKGATAAPQRKS
EDDSAVPLAKAAPKSGPSVPVSVQTKDDVYEAFMKEMEGLL
Sequence of entity 10 (Z), FASTA
>9R3D_10 pre-mRNA (chains Z)
GCCGCCAAGGUAUGUAUCAAGCUUACAAG
Sequence of entity 11 (q), FASTA
>9R3D_11 Ubiquitin-like protein 5 (chains q)
MIEVVCNDRLGKKVRVKCNTDDTIGDLKKLIAAQTGTRWNKIVLKKWYTIFKDHVSLGDY
EIHDGMNLELYYQ
Sequence of entity 12 (r), FASTA
>9R3D_12 Zinc finger matrin-type protein 2 (chains r)
MASGSGTKNLDFRRKWDKDEYEKLAEKRLTEEREKKDGKPVQPVKRELLRHRDYKVDLES
KLGKTIVITKTTPQSEMGGYYCNVCDCVVKDSINFLDHINGKKHQRNLGMSMRVERSTLD
QVKKRFEVNKKKMEEKQKDYDFEERMKELREEEEKAKAYKKEKQKEKKRRAEEDLTFEED
DEMAAVMGFSGFGSTKKSY
Ligands and cofactors
| ID | Name | Formula | Copies |
|---|
| IHP | Inositol hexakisphosphate | C6 H18 O24 P6 | 1 |
Primary citation
cryo-EM structure of the human pre-Bact-OTS complex. Zhang, Z., Kumar, V., Dybkov, O. et al. To be published.
Other PDB entries of the same protein (UniProt O75643 (AlphaFold model), which also has an AlphaFold model), best resolution first:
- 2Q0Z 2.0 Å, Crystal structure of Q9P172/Sec63 from Homo sapiens. Northeast Structural Genomics…
- 8BCE 2.05 Å, Human Brr2 Helicase Region in complex with C-tail deleted Jab1 and compound 76
- 8BC9 2.3 Å, Human Brr2 Helicase Region in complex with C-tail deleted Jab1 and compound 24
- 8BCC 2.35 Å, Human Brr2 Helicase Region in complex with C-tail deleted Jab1 and compound 39
- 8BCB 2.38 Å, Human Brr2 Helicase Region in complex with C-tail deleted Jab1 and compound 34
- 8BC8 2.39 Å, Human Brr2 Helicase Region in complex with C-tail deleted Jab1 and compound 18
- 8BCG 2.39 Å, Human Brr2 Helicase Region in complex with C-tail deleted Jab1 and compound 86
- 6S8Q 2.39 Å, Human Brr2 Helicase Region in complex with C-tail deleted Jab1
- 9S2E 2.4 Å, Human intron-lariat spliceosome-Aquarius (ILSaqr)
- 8BCF 2.42 Å, Human Brr2 Helicase Region in complex with C-tail deleted Jab1 and compound 78
- 7BDK 2.52 Å, Human Brr2 Helicase Region in complex with C-tail deleted Jab1 and ADP
- 7BDJ 2.59 Å, Human Brr2 Helicase Region in complex with C-tail deleted Jab1 and mant-ATPgammaS
Browse structure collections
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