9UJS: DNA-directed RNA polymerase subunit

RNA polymerase II elongation complex stalled at SHL(-4) of the H3-H4 octasome. Determined by electron microscopy at 3.62 Å resolution. Released 15 Apr 2026.

Method: Electron microscopy
Resolution: 3.62 Å
Organisms: Komagataella phaffii GS115, synthetic construct, Homo sapiens
Chains: 23
Atoms: 42,216
Mol. weight: 745.69 kDa
Released: 15 Apr 2026

Explore 9UJS in 3D Show helices and sheets RCSB PDB PDBe

Secondary structure: helices and β-sheets

9UJS contains 210 α-helices and 238 β-strands across 20 chains. Residue ranges use author residue numbering, as in the PDB file, from PDBe. To see them in 3D, choose the Cartoon representation with Secondary structure coloring: helices, sheets and coils get different colors.

Chain a: 3 helices, 1 β-strand

Element	Residues	Length	Sheet
α-helix	64-78	15
α-helix	86-113	28
β-strand	118-119	2	78
α-helix	122-130	9

Chain A: 79 helices, 67 β-strands

Element	Residues	Length	Sheet
α-helix	10	1
β-strand	11	1	1
β-strand	16-18	3	2
β-strand	21	1	3
α-helix	22-23	2
α-helix	24-30	7
β-strand	34	1	4
α-helix	48-49	2
α-helix	56-58	3
β-strand	82-91	10	4
α-helix	93-95	3
α-helix	96-105	10
β-strand	106	1	5
β-strand	113	1	5
α-helix	120-125	6
α-helix	131-143	13
β-strand	147	1	6
α-helix	170	1
β-strand	171	1	6
α-helix	172-173	2
β-strand	174-178	5	7
β-strand	181-186	6	7
β-strand	199-202	4	7
α-helix	205-212	8
α-helix	217-222	6
β-strand	226	1	8
β-strand	230	1	8
α-helix	232-235	4
β-strand	236-242	7	4
α-helix	245-247	3
β-strand	251-252	2	9
β-strand	258-259	2	9
α-helix	260-261	2
α-helix	262-281	20
α-helix	289-305	17
α-helix	313-315	3
β-strand	316	1	10
β-strand	322	1	10
α-helix	326-330	5
α-helix	336-340	5
β-strand	344-345	2	11
β-strand	349-353	5	12
β-strand	354-356	3	13
β-strand	364-368	5	14
α-helix	369-374	6
β-strand	376-380	5	15
α-helix	383-385	3
α-helix	386-395	10
β-strand	403-407	5	15
α-helix	422-424	3
β-strand	432-436	5	15
β-strand	443	1	16
β-strand	446	1	12
α-helix	453-455	3
β-strand	459	1	16
β-strand	460-464	5	14
β-strand	469-471	3	13
α-helix	473-475	3
α-helix	476-479	4
β-strand	487-489	3	12
α-helix	496-501	6
α-helix	502-506	5
α-helix	508-511	4
β-strand	513-514	2	17
β-strand	519-520	2	17
α-helix	526-535	10
β-strand	541-543	3	18
α-helix	544-552	9
α-helix	561-564	4
β-strand	566-568	3	18
β-strand	572-574	3	18
α-helix	575-582	8
α-helix	584	1
β-strand	588-591	4	19
β-strand	606-609	4	19
β-strand	613-615	3	19
α-helix	620-623	4
α-helix	630-661	32
α-helix	667-670	4
α-helix	674-699	26
α-helix	703-706	4
α-helix	711-737	27
α-helix	743-750	8
α-helix	756-759	4
α-helix	760-764	5
β-strand	767-768	2	20
β-strand	771	1	21
β-strand	774	1	21
α-helix	795-797	3
β-strand	800-801	2	20
α-helix	811-845	35
β-strand	850	1	22
β-strand	858	1	22
β-strand	864	1	22
α-helix	869-871	3
β-strand	874	1	23
β-strand	879-883	5	24
α-helix	891-898	8
α-helix	909-910	2
α-helix	911-913	3
β-strand	914	1	25
α-helix	917-919	3
α-helix	927-943	17
α-helix	944-948	5
β-strand	955-959	5	24
α-helix	962-973	12
α-helix	979-980	2
β-strand	981	1	25
α-helix	985-995	11
α-helix	1007-1016	10
α-helix	1018-1027	10
α-helix	1030-1031	2
α-helix	1032-1037	6
α-helix	1041-1056	16
β-strand	1059	1	23
α-helix	1066-1079	14
α-helix	1100-1107	8
β-strand	1117-1120	4	26
β-strand	1122	1	27
α-helix	1124-1127	4
α-helix	1131-1140	10
β-strand	1144	1	28
β-strand	1149-1156	8	29
α-helix	1169-1176	8
β-strand	1193-1199	7	29
α-helix	1201-1207	7
α-helix	1211-1221	11
β-strand	1226-1228	3	29
β-strand	1238-1244	7	29
α-helix	1261-1273	13
β-strand	1275	1	28
β-strand	1284-1295	12	26
β-strand	1301-1312	12	26
α-helix	1316-1319	4
β-strand	1325	1	27
β-strand	1331-1332	2	26
α-helix	1335-1359	25
α-helix	1360-1362	3
α-helix	1368-1378	11
α-helix	1399-1403	5
α-helix	1408-1417	10
β-strand	1421-1422	2	2
α-helix	1427-1432	6
α-helix	1440-1442	3
β-strand	1445-1448	4	30
α-helix	1450-1455	6

Chain b: 4 helices, 1 β-strand

Element	Residues	Length	Sheet
α-helix	25-27	3
α-helix	31-41	11
β-strand	45-46	2	78
α-helix	50-75	26
α-helix	83-93	11

Chain B: 50 helices, 80 β-strands

Element	Residues	Length	Sheet
α-helix	14-27	14
α-helix	32-40	9
α-helix	41-45	5
α-helix	46-50	5
β-strand	55-58	4	31
β-strand	71	1	32
β-strand	74-84	11	31
β-strand	88-90	3	33
α-helix	91	1
β-strand	96-98	3	33
α-helix	101-106	6
β-strand	112-124	13	31
β-strand	127	1	32
β-strand	155-162	8	31
α-helix	177-182	6
β-strand	194-196	3	34
β-strand	199-203	5	34
β-strand	205-209	5	35
α-helix	210	1
β-strand	214-218	5	36
β-strand	225-234	10	36
β-strand	241-250	10	36
β-strand	261-264	4	36
β-strand	272-273	2	36
α-helix	274-280	7
α-helix	286-293	8
α-helix	300-313	14
α-helix	319-329	11
α-helix	332-333	2
α-helix	338-350	13
α-helix	364-382	19
α-helix	387-390	4
α-helix	394-396	3
β-strand	397-400	4	35
α-helix	402-430	29
α-helix	444-455	12
α-helix	462-464	3
β-strand	473-475	3	34
α-helix	481-487	7
β-strand	490	1	37
β-strand	491-492	2	35
α-helix	503-505	3
α-helix	509-511	3
β-strand	515	1	38
β-strand	531	1	37
β-strand	532	1	38
β-strand	537-538	2	39
α-helix	545-553	9
α-helix	556	1
β-strand	557-558	2	40
α-helix	559-561	3
β-strand	571-575	5	40
β-strand	578-583	6	40
α-helix	586-598	13
β-strand	607-610	4	40
β-strand	616-620	5	40
β-strand	626-632	7	39
β-strand	633	1	41
β-strand	645	1	41
α-helix	649-657	9
α-helix	678-684	7
β-strand	687-691	5	39
α-helix	694-697	4
β-strand	700-701	2	39
α-helix	704-707	4
β-strand	740-741	2	39
α-helix	745-748	4
α-helix	764-773	10
α-helix	774-776	3
β-strand	777	1	42
β-strand	779	1	43
β-strand	793-796	4	43
β-strand	800	1	44
β-strand	804-805	2	45
β-strand	821-825	5	46
β-strand	839-842	4	46
α-helix	843-847	5
β-strand	853-862	10	43
β-strand	865-867	3	47
β-strand	870-871	2	47
β-strand	873-874	2	48
β-strand	882-883	2	48
β-strand	898	1	49
α-helix	899-900	2
β-strand	904-905	2	43
β-strand	910-912	3	49
β-strand	914-917	4	48
α-helix	918-919	2
β-strand	934-936	3	48
β-strand	939-940	2	49
α-helix	941-942	2
β-strand	947-956	10	43
β-strand	962-972	11	43
β-strand	979-982	4	46
β-strand	987-994	8	46
β-strand	1001	1	50
β-strand	1010-1012	3	46
α-helix	1016-1020	5
α-helix	1023-1038	16
α-helix	1041	1
β-strand	1042-1043	2	45
α-helix	1052-1061	10
β-strand	1069-1070	2	51
β-strand	1073	1	50
α-helix	1079	1
β-strand	1080	1	50
α-helix	1081	1
β-strand	1085-1086	2	51
β-strand	1088-1094	7	46
β-strand	1095	1	42
α-helix	1099-1101	3
β-strand	1104-1106	3	12
β-strand	1111	1	52
α-helix	1117	1
β-strand	1118	1	52
α-helix	1119	1
β-strand	1129-1130	2	11
α-helix	1132-1141	10
α-helix	1144-1152	9
β-strand	1157-1158	2	53
β-strand	1161-1163	3	1
β-strand	1172-1174	3	54
β-strand	1179-1181	3	54
β-strand	1182	1	55
β-strand	1187	1	55
β-strand	1191-1193	3	1
β-strand	1196-1197	2	53
α-helix	1198-1209	12
β-strand	1212	1	3
β-strand	1215-1217	3	2

Chains c and e: 3 helices, 1 β-strand

Element	Residues	Length	Sheet
α-helix	64-78	15
α-helix	86-113	28
β-strand	119	1	79
α-helix	121-131	11

Chain C: 11 helices, 12 β-strands

Element	Residues	Length	Sheet
β-strand	6-12	7	56
β-strand	17-22	6	56
α-helix	26-38	13
β-strand	42-53	12	57
α-helix	59-67	9
β-strand	71	1	58
α-helix	80-81	2
β-strand	96-103	8	57
β-strand	111-114	4	59
α-helix	115-117	3
β-strand	119-120	2	57
α-helix	131	1
β-strand	132	1	58
α-helix	133	1
β-strand	143-147	5	59
β-strand	152-162	11	57
α-helix	168-170	3
β-strand	176-180	5	56
α-helix	197-200	4
α-helix	202-204	3
α-helix	211-214	4
β-strand	227-232	6	56
α-helix	240-265	26

Chain d: 4 helices, 1 β-strand

Element	Residues	Length	Sheet
α-helix	26-28	3
α-helix	31-40	10
β-strand	46	1	79
α-helix	50-76	27
α-helix	83-94	12

Chain D: 9 helices, 5 β-strands

Element	Residues	Length	Sheet
β-strand	4-6	3	30
β-strand	24	1	60
β-strand	29	1	60
α-helix	32-34	3
β-strand	37-39	3	61
β-strand	45-47	3	61
α-helix	53-71	19
α-helix	86-90	5
α-helix	96-110	15
α-helix	118-128	11
α-helix	139-147	9
α-helix	153-159	7
α-helix	161-163	3
α-helix	171-181	11

11 more chain groups are not listed. Open the entry in the viewer and use the sequence panel to see them.

Molecules and chains

Molecule	Chains	Type	Length	Organism	UniProt
DNA-directed RNA polymerase subunit	A	protein	1743	Komagataella phaffii GS115	C4R4Y0
DNA-directed RNA polymerase subunit beta	B	protein	1227	Komagataella phaffii GS115	C4QZQ7 (AlphaFold model)
RNA polymerase II third largest subunit B44, part of central core	C	protein	304	Komagataella phaffii GS115	C4R7L2 (AlphaFold model)
RNA polymerase II subunit B32	D	protein	186	Komagataella phaffii GS115	C4R2U9 (AlphaFold model)
RNA polymerase subunit ABC27, common to RNA polymerases I, II, and III	E	protein	214	Komagataella phaffii GS115	C4R3P8
RNA polymerase subunit ABC23, common to RNA polymerases I, II, and III	F	protein	155	Komagataella phaffii GS115	C4R1V1
RNA polymerase II subunit	G	protein	171	Komagataella phaffii GS115	C4R9A1
DNA-directed RNA polymerases I, II, and III subunit RPABC3	H	protein	145	Komagataella phaffii GS115	C4R273
DNA-directed RNA polymerase subunit	I	protein	115	Komagataella phaffii GS115	F2QPE6
RNA polymerase subunit ABC10-beta, common to RNA polymerases I, II, and III	J	protein	72	Komagataella phaffii GS115	C4R009
RNA polymerase II subunit B12.5	K	protein	118	Komagataella phaffii GS115	C4R3Z5
RNA polymerase subunit ABC10-alpha	L	protein	72	Komagataella phaffii GS115	F2QMI1

5 more molecules are not listed.

Sequence of entity 1 (A), FASTA

>9UJS_1 DNA-directed RNA polymerase subunit (chains A)
MSQFPYSSAPLRSVKEVQFGLLSPEEIRAISVVKIEYPEIMDESRQRPREGGLNDPKLGS
IDRNFKCQTCGEGMAECPGHFGHMELAKPVFHIGFIPKIKKVCECICMNCGKLLLDETNP
TMAQAIRIRDPKKRFNAVWQLCKTKMVCEADAPVDEYSEQKVVSRGGCGNTQPVVRKDGM
KLWGTWKKSGFSDRDAQPERKLLTPGEILNVFKHISPEDCFRLGFNEDYARPEWMIITVL
PVPPPQVRPSIAMDETTQGQDDLTHKLSDILKANINVQKLEMDGSPQHIINEVEQLLQFH
VATYMDNDIAGQPQALQKSGRPVKAIRARLKGKEGRLRGNLMGKRVDFSARTVISGDPNL
ELDQVGVPISIAKTLSYPETVTQYNIHRLTEYVRNGPNEHPGAKYVIRDNGDRIDLRYHK
RAGDIVLQYGWKVERHLMDDDPVLFNRQPSLHKMSMMAHRVKVMPYSTFRLNLSVTSPYN
ADFDGDEMNLHVPQSEETRAELSQLCAVPLQIVSPQSNKPVMGIVQDTLCGVRKMTLRDT
FIEYEQVMNMLFWVPSWDGVVPQPAILKPKPLWTGKQLLSIAIPSGIHLQRTDGGNSLLS
PKDNGMLIVDGKVMFGVVDKKTVGSGGGGLIHTVMREKGPKICAELFGNIQKVVNYWLLH
NGFSIGIGDAIADASTMKEITHAISSAKEQVQEIIYKAQHNELELKPGMTLRESFEGEVS
RTLNDARDSAGRSAEMNLKDLNNVKQMVSAGSKGSFINIAQMSACVGQQMVEGKRIAFGF
ADRSLPHFTKDDFSPESKGFVENSYLRGLTPQEFFFHAMAGREGLIDTAVKTAETGYIQR
RLVKALEDIMVHYDGTTRNSLGDIIQFLYGEDGLDGTQVERQTIDTIPGSDKAFHKRYYV
DLMDEKNSIKPDVIEYAADILGDVELQKELNSEYEQLVSDRKFLREIVFVNGDHNWPLPV
NLRRIIQNAQQIFHLDRAKASDLTIPEIIHGVRDLCKKLFVLRGENELIKEAQQNATSLF
QCLVRARLATRRILEEFRLNRDAFEWVLGTIEAQFQRSLVHPGEMVGVIAAQSIGEPATQ
MTLNTFHYAGVSSKNVTLGVPRLKEILNVAKNIKTPALTVYLDREIALDIEKAKVIQSSI
EYTTLKNVTSATEIYYDPDPTSTVIEEDFDTVEAYFSIPDEKVEETIDKQSPWLLRLELD
RARMLDKQLTMNQVADKISEVFSDDLFVMWSEDNADKLIIRCRVIRDPKAMDEELEAEED
QMLKRIEAHMLDLIALRGIPGISKVYMVKHKVSVPDESGEYKNEELWALETDGINLAEVM
AVPGVDSSRTYSNSFVEILSVLGIEATRSSLYKEILNVIAFDGSYVNYRHMALLVDVMTS
RGYLMAITRHGINRADTGALMRCSFEETVEILFEAGAAAELDDCRGVSENVMLGQLAPMG
TGAFDVMIDEKLLTSLPADYAPTMPLFKGKATQGSATPYDNNAQYDDEFNHDDVADVMFS
PMAETGSGDDRSGGLTEYAGIQSPYQPTSPGLSATSPGFAPTSPGFAPTSPRYSPTSPGY
SPTSPSYSPTSPSYSPTSPSYSPTSPSYSPTSPSYSPTSPSYSPTSPSYSPTSPSYSPTS
PSYSPTSPQYSPTSPQYSPTSPQYSPTSPQYSPTSPQYSPTSPQYSPTSPQYSPTSPQYS
PTSPQYSPTSPQYSPTSPQYSPTSPQYSPTSPQYSPTSPQYSPASPQYSPSRHSPNGESK
EGE

Sequence of entity 2 (B), FASTA

>9UJS_2 DNA-directed RNA polymerase subunit beta (chains B)
MSYDPYSIDDTITTEDCWTVISAFFEEKGLVSQQLDSFDEFMETSIQDLVWEEPRLILDQ
PAQHTNEKDNINKRYEIRFGKIYLSRPTMTEADGTTHAMFPQEARLRNLTYSSPVYLDME
KSMFTSIDDEGNPNATLDWQQVHEPIKDGVEEGNKVHIGKVPIMLRSKFCSLRTLDEVDL
YKMKECPYDMGGYFVINGSEKVLIAQERSAANIVQVFKKAAPSPISHVAEIRSALEKGSR
LISTMQIKLYGREDKGTGRTIKATLPYVKQDIPIVIVFRALGVVPDGEILQHICYDENDW
QMLEMLKPCIEEGFVIQDKEVALDFIGRRGSAALGIRREKRIQYAKDILQKELLPHITQE
EGFETRKTFFLGYMVNRLLLCALERKDQDDRDHFGKKRLDLAGPLLANLFRILFRKLTRE
IYRYMQRCIETDRDFNLNLAVKSTTITSGLKYSLATGNWGEQKKAMSSRAGVSQVLNRYT
YSSTLSHLRRTNTPIGRDGKLAKPRQLHNTHWGLVCPAETPEGQACGLVKNLSLLSGISI
GSPSEPIINFLEEWGMEPLEDYDPAQHTKSTRIFVNGVWTGIHRDPSMLVSTMRDLRRSG
AISPEVSIIRDIREREFKIFTDVGRVYRPLFIVEDDESKDNKGELRITKEHIRKIQQGYD
DDAMNDDSEEQEQDVYGWSSLVTSGVIEYVDGEEEETIMIAMTPEDLQTRSLEQKEIDLN
DTAKRIKPEMSTSSHHTFTHCEIHPSMILGVAASIIPFPDHNQSPRNTYQSAMGKQAMGV
FLTNYNVRMDTMANILYYPQKPLAKTQAMEYLKFRELPAGQNAIVAIACYSGYNQEDSMI
MNQSSIDRGLFRSLFFRSYMDQEKRFGISIVEEFEKPTRATTLRLKHGTYEKLDEDGLIA
PGVRVSGDDIIIGKTTPIPPDTEELGQRTKYHTKRDASTPLRSTENGIVDQVLLTTNQEG
LKFVKVRMRTTKVPQIGDKFASRHGQKGTIGVTYRHEDMPFSAEGIVPDLIINPHAIPSR
MTVAHLIECLLSKVGSIRGYEGDATPFTDLTVDAVSNLLRDNGYQSRGFEVMYNGHTGKK
LMAQVFFGPTYYQRLRHMVDDKIHARARGPVQVLTRQPVEGRSRDGGLRFGEMERDCMIA
HGAAGFLKERLMEASDAFRVHVCGICGLMSVIANLKKNQFECRSCKNKTNIYQLHIPYAA
KLLFQELMAMNIAPRLYTERSGVSMRS

Sequence of entity 3 (C), FASTA

>9UJS_3 RNA polymerase II third largest subunit B44, part of central core (chains C)
MSKEPKVNIINAQDDEVELMLSDVNLSLANSLRRTMLAEVPTLAIDLVEIKMNTSVLADE
FISHRLGLIPLVSEDVEEMKYSRDCTCEDYCDECSVVLELSARHEGEEGTTDVYSSSLIK
VSGPGNLNVGEPVRRDDYDQGILLCKLRNHQELNIRCIAKKGIAKEHAKWSPCSAIAFEY
DPHNKLKHTDFWFEVDAKKEWPDSKYATWEEPPKPGEVFDYKAKPNRFYMTVETTGSLKA
NQVFSRGIKTLQEKLANVLFELENSRPANTTAYGGATAYGGQTVYGRETSYGGNTNYGDY
NAPY

Sequence of entity 4 (D), FASTA

>9UJS_4 RNA polymerase II subunit B32 (chains D)
MNVSTSTVGARRRRAKQQVDDEENATLLRLGPEFALKQYDHDGNEHDLIALSLSESRLLI
REALKARSRARNGGVDIESSNGEIDDDELAKVTSGAVANGVVKKTLDYLNTFARFKDEET
CTAVDQLLHNSSDCSVLHPFEIAQLSSLGCEDVDEAITLIPSLAAKKEVNLQRILDELNR
LEDPYK

Sequence of entity 5 (E), FASTA

>9UJS_5 RNA polymerase subunit ABC27, common to RNA polymerases I, II, and III (chains E)
MEDNNRIISRLWRSFRTVKEMAADRGYFISQEEMDQSLEEFRSKICDSMGNPQRKLMSFL
ANPTPEALEKYSDLGTLWVEFCDEPSVGIKTMRNFCLRIQEKNFSTGIFIYQNNITPSAN
KMIPTVSPAIIETFQESDLVVNITHHELVPKHIRLSDGEKSQLLQRYKLKESQLPRIQRE
DPVARYLGLKRGQVVKIIRRSETSGRYASYRICL

Sequence of entity 6 (F), FASTA

>9UJS_6 RNA polymerase subunit ABC23, common to RNA polymerases I, II, and III (chains F)
MSEDEAFNEQTENFENFEDEHFSDDNFEDRSTQPEDYAVGVTADGRQIINGDGIQEVNGT
IKAHRKRSNKELAILKEERTTTPYLTKYERARILGTRALQISMNAPVLVDIEGETDPLQI
AMKELSQRKIPLVIRRYLPDGSYEDWGCDELIVDN

Sequence of entity 7 (G), FASTA

>9UJS_7 RNA polymerase II subunit (chains G)
MFFLKDLSLILTLHPSYFGPQMNQYLREKLLTDVEGTCTGQFGYIVTVLDGMNIDVGKGR
IIPGSGSAEFEVKYRAVVWKPFKGEVVDAIVSNVSPIGFFADVGPLNVFVSTRLIPDNLV
YNPSNSPPAYMSNDELITKGSKVRLKVVGTRTDVNEIYAIGSIKEDFLGAI

Sequence of entity 8 (H), FASTA

>9UJS_8 DNA-directed RNA polymerases I, II, and III subunit RPABC3 (chains H)
MSSALFDDIFTVQTVDNGRYNKVSRIIGISTTNSAIKLTLDINNEMFPVSQDDSLTVTLA
NSLSLDGEDESANFSKSWRPPKPTDKSLADDYDYVMFGTVYKFEEGDEDKIKVYVSFGGL
LMCLEGGYKSLASLKQDNLYILIRR

Sequence of entity 9 (I), FASTA

>9UJS_9 DNA-directed RNA polymerase subunit (chains I)
MASFRFCLECNNMLYPKEDKENQRLLYSCRNCDYTELAEDPKVYRHELITNIGETAGIVD
DIGQDPTLPRSDKECPECHSRDCVFFQSQQRRKDTNMTLFYVCLNCKKTFRDESE

Sequence of entity 10 (J), FASTA

>9UJS_10 RNA polymerase subunit ABC10-beta, common to RNA polymerases I, II, and III (chains J)
MIIPVRCFSCGKVVGDKWDAYLRLLEEGKQEGDALDELKLKRYCCRRMVLTHVDLIEKFL
RYNPLEKKDFDS

Sequence of entity 11 (K), FASTA

>9UJS_11 RNA polymerase II subunit B12.5 (chains K)
MNAPDRFELFILPDDVPKLKITPDSRVPNCIIIKFEREDHTLANLLREELALYPDVTFVA
YKVEHPLFANFVMRLQTEEGTRPKQALERACASIINKLKTLDHKFNEEWNIKNFSLND

Sequence of entity 12 (L), FASTA

>9UJS_12 RNA polymerase subunit ABC10-alpha (chains L)
MSREGFVAPSGTDLAAAASGVAPNKHYGVKYTCGACAHNFSLNKSDPVRCKECGHRVIYK
ARTKRMIQFDAR

Primary citation

Structural basis of RNA polymerase II transcription on the histone H3-H4 octasome. Ho, C.H., Nozawa, K., Nishimura, M. et al. J Biol Chem (2026) 302:111340-111340. DOI 10.1016/j.jbc.2026.111340 · PubMed

Other PDB entries of the same protein (UniProt C4R4Y0), best resolution first:

5XOG 3.0 Å, RNA Polymerase II elongation complex bound with Spt5 KOW5 and Elf1
7XN7 3.1 Å, RNA polymerase II elongation complex containing Spt4/5, Elf1, Spt6, Spn1 and Paf1C
8JH4 3.2 Å, RNA polymerase II elongation complex containing 60 bp upstream DNA loop, stalled at…
8YFQ 3.3 Å, Cryo EM structure of Komagataella phaffii RNAPII-Rat1-Rai1 pre-termination complex
7XSZ 3.4 Å, RNA polymerase II elongation complex transcribing a nucleosome (EC115)
8YFR 3.4 Å, Cryo EM structure of Komagataella phaffii Rat1-Rai1 complex bound within the RNAPII cleft
9WMS 3.4 Å, Co-transcriptional histone H3K36 methylation complex containing RNA polymerase II…
9II7 3.5 Å, RNA polymerase II elongation complex stalled at SHL(-1) of the nucleosome containing…
9WMT 3.59 Å, Co-transcriptional histone H3K36 methylation complex containing RNA polymerase II…
7XSE 3.6 Å, RNA polymerase II elongation complex transcribing a nucleosome (EC42)
9WMW 3.62 Å, Co-transcriptional histone H3K36 methylation complex containing RNA polymerase II…
8JH3 3.7 Å, RNA polymerase II elongation complex containing 40 bp upstream DNA loop, stalled at…

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