9XRL: Mouse cytoplasmic lattice (CPL) repeating unit

Structure of mouse cytoplasmic lattice (CPL) repeating unit. Determined by electron microscopy at 3.74 Å resolution. Released 18 Mar 2026.

Method: Electron microscopy
Resolution: 3.74 Å
Organism: Mus musculus
Chains: 73
Atoms: 281,032
Mol. weight: 4735.88 kDa
Ligands: ADP, MG, GTP, ZN
Released: 18 Mar 2026

Explore 9XRL in 3D Show helices and sheets RCSB PDB PDBe

Secondary structure: helices and β-sheets

9XRL contains 1,748 α-helices and 2,042 β-strands across 73 chains. Residue ranges use author residue numbering, as in the PDB file, from PDBe. To see them in 3D, choose the Cartoon representation with Secondary structure coloring: helices, sheets and coils get different colors.

Chain 0: 49 helices, 29 β-strands

Element	Residues	Length	Sheet
α-helix	98-109	12
α-helix	120-127	8
β-strand	140-145	6	528
α-helix	151-164	14
β-strand	173-178	6	528
β-strand	187-188	2	547
α-helix	190-197	8
α-helix	202-208	7
α-helix	212-214	3
β-strand	215-220	6	528
α-helix	222-227	6
β-strand	244-245	2	547
α-helix	246-254	9
β-strand	263-269	7	528
α-helix	270-272	3
α-helix	273-279	7
β-strand	284-288	5	528
α-helix	293-303	11
α-helix	310-318	9
α-helix	321-326	6
α-helix	330-346	17
α-helix	347-349	3
α-helix	357-370	14
α-helix	374	1
α-helix	381-399	19
β-strand	405-406	2	548
α-helix	407-412	6
α-helix	417-423	7
β-strand	429-432	4	548
β-strand	439-442	4	548
α-helix	445-458	14
α-helix	464-467	4
α-helix	486-488	3
α-helix	489-497	9
α-helix	501-511	11
α-helix	519-531	13
α-helix	539-552	14
α-helix	555-561	7
β-strand	567-571	5	549
α-helix	575-585	11
β-strand	593-597	5	549
α-helix	624-636	13
β-strand	644-646	3	549
β-strand	651	1	550
α-helix	654-664	11
β-strand	673-675	3	549
β-strand	680	1	550
α-helix	682-687	6
α-helix	690-694	5
β-strand	700-702	3	549
α-helix	710-721	12
β-strand	729-731	3	549
α-helix	739-750	12
β-strand	757-759	3	549
α-helix	767-777	11
β-strand	786-788	3	549
α-helix	796-806	11
β-strand	814-816	3	549
α-helix	823-835	13
β-strand	843-845	3	549
α-helix	854-863	10
β-strand	871-873	3	549
α-helix	880-891	12
β-strand	900-902	3	549
α-helix	910-912	3
α-helix	913-922	10
β-strand	928-930	3	549
α-helix	937-948	12
β-strand	957-959	3	549
α-helix	967-969	3
α-helix	970-979	10
β-strand	985-987	3	549
α-helix	995-1005	11
β-strand	1014-1016	3	549
α-helix	1019-1021	3
α-helix	1024-1036	13
β-strand	1041-1042	2	549
α-helix	1046-1048	3

Chain 1: 4 helices, 3 β-strands

Element	Residues	Length	Sheet
α-helix	36-39	4
β-strand	42-43	2	1
α-helix	48-49	2
β-strand	57-60	4	1
β-strand	66-68	3	1
α-helix	73-83	11
α-helix	84-88	5

Chain 2: 49 helices, 29 β-strands

Element	Residues	Length	Sheet
α-helix	98-109	12
α-helix	120-127	8
β-strand	140-145	6	311
α-helix	151-164	14
β-strand	173-178	6	311
β-strand	187-188	2	312
α-helix	190-197	8
α-helix	202-208	7
α-helix	212-214	3
β-strand	215-220	6	311
α-helix	222-227	6
α-helix	235-237	3
β-strand	244-245	2	312
α-helix	246-254	9
β-strand	263-269	7	311
α-helix	273-276	4
β-strand	284-288	5	311
α-helix	293-303	11
α-helix	311-318	8
α-helix	321-326	6
α-helix	330-346	17
α-helix	347-349	3
α-helix	357-370	14
α-helix	374	1
α-helix	381-399	19
β-strand	405-406	2	313
α-helix	407-412	6
α-helix	417-423	7
β-strand	429-432	4	313
β-strand	439-442	4	313
α-helix	445-458	14
α-helix	464-467	4
α-helix	486-488	3
α-helix	489-497	9
α-helix	501-511	11
α-helix	519-531	13
α-helix	539-552	14
α-helix	555-561	7
β-strand	567-571	5	314
α-helix	575-585	11
β-strand	593-597	5	314
α-helix	624-636	13
β-strand	644-646	3	314
β-strand	651	1	315
α-helix	654-664	11
β-strand	673-675	3	314
β-strand	680	1	315
α-helix	682-684	3
α-helix	686-694	9
β-strand	700-702	3	314
α-helix	710-721	12
β-strand	729-731	3	314
α-helix	739-750	12
β-strand	757-759	3	314
α-helix	767-777	11
β-strand	786-788	3	314
α-helix	796-806	11
β-strand	814-816	3	314
α-helix	823-835	13
β-strand	843-845	3	314
α-helix	854-863	10
β-strand	871-873	3	314
α-helix	880-891	12
β-strand	900-902	3	314
α-helix	910-912	3
α-helix	913-922	10
β-strand	928-930	3	314
α-helix	937-948	12
β-strand	957-959	3	314
α-helix	967-969	3
α-helix	970-979	10
β-strand	985-987	3	314
α-helix	995-1005	11
β-strand	1014-1016	3	314
α-helix	1019-1021	3
α-helix	1024-1036	13
β-strand	1041-1042	2	314
α-helix	1046-1048	3

Chains 3 and 9: 5 helices, 3 β-strands

Element	Residues	Length	Sheet
α-helix	36-39	4
β-strand	42-43	2	33
α-helix	48-49	2
α-helix	52-54	3
β-strand	57-60	4	33
β-strand	66-68	3	33
α-helix	73-83	11
α-helix	84-88	5

Chain 4: 7 helices, 5 β-strands

Element	Residues	Length	Sheet
β-strand	13-17	5	58
β-strand	20-23	4	58
α-helix	35-38	4
β-strand	42-46	5	59
α-helix	48-50	3
α-helix	51-55	5
α-helix	57-59	3
α-helix	62-69	8
β-strand	72-76	5	59
α-helix	83	1
β-strand	84-89	6	59
α-helix	92-123	32

Chain 5: 48 helices, 29 β-strands

Element	Residues	Length	Sheet
α-helix	98-109	12
α-helix	120-127	8
β-strand	140-145	6	172
α-helix	151-164	14
β-strand	173-178	6	172
β-strand	187-188	2	173
α-helix	190-197	8
α-helix	202-208	7
α-helix	212-214	3
β-strand	215-220	6	172
α-helix	222-227	6
β-strand	244-245	2	173
α-helix	246-254	9
β-strand	263-269	7	172
α-helix	273-279	7
β-strand	284-288	5	172
α-helix	293-303	11
α-helix	311-318	8
α-helix	321-326	6
α-helix	330-346	17
α-helix	347-349	3
α-helix	357-370	14
α-helix	374	1
α-helix	381-399	19
β-strand	405-406	2	174
α-helix	407-412	6
α-helix	417-423	7
β-strand	429-432	4	174
β-strand	439-442	4	174
α-helix	445-458	14
α-helix	464-467	4
α-helix	486-488	3
α-helix	489-497	9
α-helix	501-511	11
α-helix	519-531	13
α-helix	539-552	14
α-helix	555-561	7
β-strand	567-571	5	175
α-helix	575-585	11
β-strand	593-597	5	175
α-helix	624-636	13
β-strand	644-646	3	175
β-strand	651	1	176
α-helix	654-664	11
β-strand	673-675	3	175
β-strand	680	1	176
α-helix	682-684	3
α-helix	686-689	4
β-strand	700-702	3	175
α-helix	710-721	12
β-strand	729-731	3	175
α-helix	739-750	12
β-strand	757-759	3	175
α-helix	767-777	11
β-strand	786-788	3	175
α-helix	796-806	11
β-strand	814-816	3	175
α-helix	823-835	13
β-strand	843-845	3	175
α-helix	854-863	10
β-strand	871-873	3	175
α-helix	880-891	12
β-strand	900-902	3	175
α-helix	910-912	3
α-helix	913-922	10
β-strand	928-930	3	175
α-helix	937-948	12
β-strand	957-959	3	175
α-helix	967-969	3
α-helix	970-979	10
β-strand	985-987	3	175
α-helix	995-1005	11
β-strand	1014-1016	3	175
α-helix	1019-1021	3
α-helix	1024-1036	13
β-strand	1041-1042	2	175
α-helix	1046-1048	3

Chain 6: 6 helices, 7 β-strands

Element	Residues	Length	Sheet
β-strand	3	1	83
β-strand	13-17	5	84
β-strand	20-23	4	84
α-helix	35-38	4
β-strand	42-46	5	85
α-helix	48-55	8
α-helix	57-59	3
α-helix	62-69	8
β-strand	72-76	5	85
β-strand	77	1	83
α-helix	83	1
β-strand	84-89	6	85
α-helix	92-123	32

Chain 7: 49 helices, 31 β-strands

Element	Residues	Length	Sheet
α-helix	98-109	12
α-helix	120-127	8
β-strand	140-145	6	150
α-helix	151-164	14
β-strand	173-178	6	150
β-strand	187-188	2	151
α-helix	190-197	8
α-helix	202-208	7
α-helix	212-214	3
β-strand	215-220	6	150
α-helix	222-227	6
β-strand	244-245	2	151
α-helix	246-254	9
β-strand	263-269	7	150
α-helix	270-272	3
α-helix	273-276	4
β-strand	284-288	5	150
α-helix	293-303	11
α-helix	310-318	9
α-helix	321-326	6
α-helix	330-346	17
α-helix	347-349	3
α-helix	357-370	14
α-helix	374	1
α-helix	381-399	19
β-strand	405-406	2	152
α-helix	407-412	6
α-helix	417-423	7
β-strand	429-432	4	152
β-strand	439-442	4	152
α-helix	445-458	14
α-helix	464-467	4
α-helix	486-488	3
α-helix	489-497	9
α-helix	501-511	11
α-helix	519-531	13
α-helix	539-552	14
α-helix	555-561	7
β-strand	567-572	6	153
α-helix	575-585	11
β-strand	593-598	6	153
β-strand	610	1	396
β-strand	611	1	397
α-helix	624-636	13
β-strand	644-646	3	153
β-strand	651	1	154
α-helix	654-664	11
β-strand	673-675	3	153
β-strand	680	1	154
α-helix	682-687	6
α-helix	688-694	7
β-strand	700-702	3	153
α-helix	710-721	12
β-strand	729-731	3	153
α-helix	739-749	11
β-strand	757-759	3	153
α-helix	767-777	11
β-strand	786-788	3	153
α-helix	796-806	11
β-strand	814-816	3	153
α-helix	823-835	13
β-strand	843-845	3	153
α-helix	854-863	10
β-strand	871-873	3	153
α-helix	880-891	12
β-strand	900-902	3	153
α-helix	910-912	3
α-helix	913-922	10
β-strand	928-930	3	153
α-helix	937-948	12
β-strand	957-959	3	153
α-helix	967-969	3
α-helix	970-979	10
β-strand	985-987	3	153
α-helix	995-1005	11
β-strand	1014-1016	3	153
α-helix	1019-1021	3
α-helix	1024-1036	13
β-strand	1041-1042	2	153
α-helix	1046-1048	3

54 more chain groups are not listed. Open the entry in the viewer and use the sequence panel to see them.

Molecules and chains

Molecule	Chains	Type	Length	Organism	UniProt
Zinc finger BED domain-containing protein 3	1, 3, 9	protein	228	Mus musculus	Q9D0L1 (AlphaFold model)
KH domain-containing protein 3	4, 6, AA	protein	440	Mus musculus	Q9CWU5 (AlphaFold model)
F-box and WD-40 domain protein 21	A	protein	468	Mus musculus	Q8BI38 (AlphaFold model)
S-phase kinase-associated protein 1	D, F, O, p, q	protein	163	Mus musculus	Q9WTX5 (AlphaFold model)
Transducin-like enhancer protein 6	8, AC, E, I, P, z	protein	581	Mus musculus	Q9WVB3
NLR family, pyrin domain containing 4F	H, Q, h, i	protein	937	Mus musculus	L7N1W9
Inactive protein-arginine deiminase type-6	B, K, S, T, U, V, W, X, Y, Z, b, c, d, f, j, m, n, o, r, s	protein	682	Mus musculus	Q8K3V4
Oocyte-expressed protein homolog	AH, AI, AJ, AT, e, g	protein	164	Mus musculus	Q9CWE6
Expressed sequence C85627	k, l	protein	469	Mus musculus	Q3TSA9
Tubulin beta-4B chain	AE, AN, t	protein	445	Mus musculus	P68372
NACHT, LRR and PYD domains-containing protein 5	0, 2, 5, 7, AB, y	protein	1059	Mus musculus	Q9R1M5
Tubulin alpha-1A chain	AF, AO, u	protein	451	Mus musculus	P68369

4 more molecules are not listed.

Sequence of entity 1 (1, 3, 9), FASTA

>9XRL_1 Zinc finger BED domain-containing protein 3 (chains 1, 3, 9)
MKSKKPLKITMEDSRRLNDPAEQGGLCPAPVGPSYSEAWGYFHLDPAQPRHRMMSAWATC
RLCGLQVGGLPNFQMWTRALCQHLSDVHLPELKKSAAPSSPTTMPCPPPPSPTMAAEGDW
ARLLEQMGELAMRGSQRELELERREAALMQAELELERKRQALKQEAQSVEQERHQLQVER
EALSKWIKKQSPGAQVPEPPSPLPLLPKEDPDIHDNNSDNDMVTKVLL

Sequence of entity 2 (4, 6, AA), FASTA

>9XRL_2 KH domain-containing protein 3 (chains 4, 6, AA)
MASLKRFQTLVPLDHKQGTLFEIIGEPKLPKWFHVECLEDPKRLYVEPRLLEIMFGKDGE
HIPHLESMLHTLIHVNVWGPERRAEIWIFGPPPFRRDVDRMLTDLAHYCRMKLMEIEALE
AGVERRRMAAHKAATQPAPVKVREAAPRPASVKVPETATQPAPVKVREAAPQPAPVQEVR
EAAPQQASVQEEVREAATEQAPVQEVREAATEQAPVQEVSEAATEQAPVQEVNEAATEQA
SVQAVREAATRPAPGKVRKAATQPAPVQVCQEATQLAPVKVREAATQPASGKVREAATQL
APVKVRKAATQLAPVKVHEAATQPAPGKVSDAATQSASVQVREAATQLSPVEATDTSQLA
QVKADEAFAQHTSGEAHQVANGQSPIEVCETATGQHSLDVSRALSQKCPEVFEWETQSCL
DGSYVIVQPPRDAWESFIIL

Sequence of entity 3 (A), FASTA

>9XRL_3 F-box and WD-40 domain protein 21 (chains A)
MEIHLPRLPLMEIFSYLDAYSLLQVAQVNKNWNELASSDVLWRKLCQKRWFYCNMVTQQL
LGKETWKEFFIYRTWQEHAKSRAKPEDFIYKEIPAEYGIQAYACYISEHGLTRNGQGRSV
ICMATSMNRISTWDIHEGVLTWVSPEQPASIKLLTTLPEMYIAVTVDMESTIKLWDCHNS
EALATNSLISPCQSLKAVITKDGPIVLIGDTLGNLNIFRIPDLYHITRLKVFPYGISELY
CSPQKKWIFLNRKHPHILPKVFYMSSLLRTSEFSAPVSTDLKFSLCQRAFWTPRREDRIT
LMSIHGPKKIKKFITFDMELEKIGNKITVKEHFFASFSLQNYEERPEWYGVSDKDVIVCS
TRFSLLLFDINGHCLQAFQYCPEQILRLWVDPLHVIVSCNDGFLDVYAWEERSQQLNKCY
RLQYSKHLPSSGLINKTLSDDVSIIQVITIRTTPCFLMAFILKSALEN

Sequence of entity 4 (D, F, O, p, q), FASTA

>9XRL_4 S-phase kinase-associated protein 1 (chains D, F, O, p, q)
MPTIKLQSSDGEIFEVDVEIAKQSVTIKTMLEDLGMDDEGDDDPVPLPNVNAAILKKVIQ
WCTHHKDDPPPPEDDENKEKRTDDIPVWDQEFLKVDQGTLFELILAANYLDIKGLLDVTC
KTVANMIKGKTPEEIRKTFNIKNDFTEEEEAQVRKENQWCEEK

Sequence of entity 5 (8, AC, E, I, P, z), FASTA

>9XRL_5 Transducin-like enhancer protein 6 (chains 8, AC, E, I, P, z)
MTSHRQSSDTFGGILPSTLSSRYLSIVNQLPEEFSSVVSEMMVHLENIFSLAENFFQAIE
RFSRTPDLLERNKMSIGVGAEGDSWPCHVSHEAPMGSAQTTENSAKEEDKQVPESAALQH
PKFKSTPGPQLPTRRRFLSESDELQDPQPVWDAEPQFCQGFLIQGLWELFMDSRQKNQQE
HGGEDSSQESKDSGLCDFKPEPQPRHRNSLSDSADPFLIKSPSALLDYYQEDVSRPQPET
QESSGRADKFLKPLSWGSEVLESSCNQPSTALWQLERFTVPQALQKVRVLKHQELLLVVA
VSSFTRHVFTCSQSGIKVWNLVNQVAEDRDPESHLKCSVQDNKVYLRTCLLSSNSRTLFA
GGYNLPGVIVWDLAAPSLYEKCQLPCEGLSCQALANTKENMALAGFTDGTVRIWDLRTQE
IVRNLKGPTNSARNLVVKDDNIWTGGLDACLRCWDLRMAKVSLEHLFQSQIMSLAHSPTE
DWLLLGLANGQHCLFNSRKRDQVLTVDTKDNTILGLKFSPNGKWWASVGMGNFITVHSMP
TGAKLFQVPEVGPVRCFDMTENGRLIITGSRDCASVYHIKY

Sequence of entity 6 (H, Q, h, i), FASTA

>9XRL_6 NLR family, pyrin domain containing 4F (chains H, Q, h, i)
MASFISDFGLIWYLRELNKKEFMKFKDFLIQEILELKLKQVSSTKVKKASREDLANLLLK
CGENQAWDMTFRILQKINRKDLTERATGAIVGNPNLYRDHLKKKLTHDCPKKFNVRIQDF
IKETFIQNDYDAFENLLISKGTERKPHMVFLKGMAGVGKTLMLKNLMLAWSKGLVFQNKF
SYAFYFCCQDVKQLKTASLAELISREWPSPSAPIEEILSQPEKLLFIIDSLEGMEWDLTK
QESELCDDCMEKQPVSTLLSSLLRRKMLPESSLLLSTTPETFEKMEDRIQCTDVKTATAF
DERSMKIYFHRLFQDRKRAQEAFSLVRENKQLFTICQVPLLCWMVATCLKEEIEKGGDPV
SLCRRTTSLYTTHIFSLFIPQSAQYPSKKSQDQLQGLCSLAAEGMWTDTFVFGKEALRRN
GIFDSDIPTLLDIGMLGKIREFENSYIFLHPSVQEVCAAIFYMLKRHVEHPSQDVKNIET
VLFMFLKKVKTQWIFLGCFIFGLLQKSEQEKLGVFFGHRLSKNIHHKLYQCLETLSGNAE
LQEQIDGMRLFSCLFEMEDEAFLVKAMNCMQQINFVAKNYSDFIVAAYCLKHCSTLKKLS
FSTENVLNEGDQSYMEELLICWNNMCSVFVRSKDIQELRIKDTNFNEPAIRVLYESLKYP
SFTLNKLVANNVSFGDNHVLFELIQNSSLQYLDLSCSFLSHNEVKLLCDILNQAECNIEK
LMIAHCKLSPDDCKIFGSILMSSKSLKVLNLASNNLNQGISSLCKALCHPHCTLEYLVLS
NCSLSEQCWDYLSEVLRQNKTLSHLDISSNDLKDEGLKILCRSLILPYCVLESLCLSCCG
ITERGCQDLAEVLKNNQNLKYLHVSYNKLKDTGVMLLCDAIKHPNCHLKDLQLEACEITD
ASNEELCYAFMQCETLQTLNLMGNAFEVSRMVFFPRF

Sequence of entity 7 (B, K, S, T, U, V, W, X, Y, Z, b, c, d, f, j, m, n, o, r, s), FASTA

>9XRL_7 Inactive protein-arginine deiminase type-6 (chains B, K, S, T, U, V, W, X, Y, Z, b, c, d, f, j, m, n, o, r, s)
MSFQNSLSLSLVNPTHALCMVGMEITLDISKCAPDKCKSFTIRGSPRILIHISSSVIAGK
EDTVVWRSMNHPTVALVRMVAPSPTVDEDKVLVSYFCPDQEVPTATAVLFLTGIEISLEA
DIYRDGQLDMPSDKQAKKKWMWGMNGWGAILLVNCSPNAVGQPDEQSFQEGPREIQNLSQ
MNVTVEGPTSILQNYQLILHTSEEEAKKTRVYWSQRGSSAYELVVGPNKPVYLLPTFENR
RKEAFYVEATEFPSPSFSGLISLSLSLVEKAHDECIPEIPLYKDTVMFRVAPYIFMPSTQ
MPLEVYLCRELQLQGFVDSVTKLSEKSKVQVVKVYEDPNRQSKWLQDEMAFCYTQAPHKT
VSLILDTPRVSKLEDFPMKYTLTPGSGYLIRQTEDHRVASLDSIGNLMVSPPVKAQGKDY
PLGRVLIGGSFYPSSEGRDMNKGLREFVYAQQVQAPVELFSDWLMTGHMDQFMCFVPTND
KNNDQKDFRLLLASPSACFELFEQKQKEGYGNVTLFEDIGAEQLLSNGRESKTISQILAD
KSFREQNTYVEKCISLNRTLLKTELGLEDKDIILIPQLFCLEQLTNVPSNQQSTKLFARP
YFPDMLQIIVLGKNLGIPKPFGPKINGTCCLEEKVCGLLEPLGLKCTFIDDFDCYLANIG
DVCASAIINRVPFAFKWWKMTP

Sequence of entity 8 (AH, AI, AJ, AT, e, g), FASTA

>9XRL_8 Oocyte-expressed protein homolog (chains AH, AI, AJ, AT, e, g)
MASHTADADAKPDSDSQKLLNVLPVSLRLRTRPWWFPIQEVSNPLVLYMEAWVAERVIGT
DQAEISEIEWMCQALLTVDSVNSGNLAEITIFGQPSAQTRMKNILLNMAAWHKENELQRA
VKVKEVEEFLKIRASSILSKLSKKGLKLAGFPLPLEGRETQMES

Sequence of entity 9 (k, l), FASTA

>9XRL_9 Expressed sequence C85627 (chains k, l)
METHLPSLPSLPMMKILSYLDAYSLLQVAQVNKNWNALASSDVLWRKLCQKRWLYCDTVI
LQLHDKETWKQFFVNRTYQEHTKTRAKPEDFTYKEICAETGIWAYACYISGRGLTRNGQG
TSVVCMLTSMTKISTWDIHEGVMTWVSPVQPTTIKLLNTLPEMHIAVTVDIHSTIKLWDC
NSSDALATNNLFFPCQTLKSVFTKDAAIVLVSDTLGNLYIFRIPDLHLISTINVFPYGIN
ELYCSPQKKWVFLSRKHPHILPKVFYMNSLLRRSEFSAPVSTVLNFSLCDKAFWTPRKED
RITLMSISAPYKVTKFVTFDMKLEEIGNQIIVTGYLIASFSLTDYEGRLECFGVSDKDVI
VCSTGSSLLLFSIYGVCLQTFDYCSEEILRLWVDPFHVIVTFIDGSLDVYAWEERCQQLS
KCYRLQNRRRLPRQSCFEKTLCDEVSIIRMVRNGRNPCYLMTYTLNIHS

Sequence of entity 10 (AE, AN, t), FASTA

>9XRL_10 Tubulin beta-4B chain (chains AE, AN, t)
MREIVHLQAGQCGNQIGAKFWEVISDEHGIDPTGTYHGDSDLQLERINVYYNEATGGKYV
PRAVLVDLEPGTMDSVRSGPFGQIFRPDNFVFGQSGAGNNWAKGHYTEGAELVDSVLDVV
RKEAESCDCLQGFQLTHSLGGGTGSGMGTLLISKIREEYPDRIMNTFSVVPSPKVSDTVV
EPYNATLSVHQLVENTDETYCIDNEALYDICFRTLKLTTPTYGDLNHLVSATMSGVTTCL
RFPGQLNADLRKLAVNMVPFPRLHFFMPGFAPLTSRGSQQYRALTVPELTQQMFDAKNMM
AACDPRHGRYLTVAAVFRGRMSMKEVDEQMLNVQNKNSSYFVEWIPNNVKTAVCDIPPRG
LKMSATFIGNSTAIQELFKRISEQFTAMFRRKAFLHWYTGEGMDEMEFTEAESNMNDLVS
EYQQYQDATAEEEGEFEEEAEEEVA

Sequence of entity 11 (0, 2, 5, 7, AB, y), FASTA

>9XRL_11 NACHT, LRR and PYD domains-containing protein 5 (chains 0, 2, 5, 7, AB, y)
MGPPEKDSKAILKARGLEEEQKSESTMSPSENVSRAILKDSGSEEVEQASERKMTSPEND
SKSIQKDQGPEQEQTSETLQSKEEDEVTEADKDNGGDLQDYKAHVIAKFDTSVDLHYDSP
EMKLLSDAFKPYQKTFQPHTIILHGRPGVGKSALARSIVLGWAQGKLFQKMSFVIFFSVR
EIKWTEKSSLAQLIAKECPDSWDLVTKIMSQPERLLFVIDGLDDMDSVLQHDDMTLSRDW
KDEQPIYILMYSLLRKALLPQSFLIITTRNTGLEKLKSMVVSPLYILVEGLSASRRSQLV
LENISNESDRIQVFHSLIENHQLFDQCQAPSVCSLVCEALQLQKKLGKRCTLPCQTLTGL
YATLVFHQLTLKRPSQSALSQEEQITLVGLCMMAAEGVWTMRSVFYDDDLKNYSLKESEI
LALFHMNILLQVGHNSEQCYVFSHLSLQDFFAALYYVLEGLEEWNQHFCFIENQRSIMEV
KRTDDTRLLGMKRFLFGLMNKDILKTLEVLFEYPVIPTVEQKLQHWVSLIAQQVNGTSPM
DTLDAFYCLFESQDEEFVGGALKRFQEVWLLINQKMDLKVSSYCLKHCQNLKAIRVDIRD
LLSVDNTLELCPVVTVQETQCKPLLMEWWGNFCSVLGSLRNLKELDLGDSILSQRAMKIL
CLELRNQSCRIQKLTFKSAEVVSGLKHLWKLLFSNQNLKYLNLGNTPMKDDDMKLACEAL
KHPKCSVETLRLDSCELTIIGYEMISTLLISTTRLKCLSLAKNRVGVKSMISLGNALSSS
MCLLQKLILDNCGLTPASCHLLVSALFSNQNLTHLCLSNNSLGTEGVQQLCQFLRNPECA
LQRLILNHCNIVDDAYGFLAMRLANNTKLTHLSLTMNPVGDGAMKLLCEALKEPTCYLQE
LELVDCQLTQNCCEDLACMITTTKHLKSLDLGNNALGDKGVITLCEGLKQSSSSLRRLGL
GACKLTSNCCEALSLAISCNPHLNSLNLVKNDFSTSGMLKLCSAFQCPVSNLGIIGLWKQ
EYYARVRRQLEEVEFVKPHVVIDGDWYASDEDDRNWWKN

Sequence of entity 12 (AF, AO, u), FASTA

>9XRL_12 Tubulin alpha-1A chain (chains AF, AO, u)
MRECISIHVGQAGVQIGNACWELYCLEHGIQPDGQMPSDKTIGGGDDSFNTFFSETGAGK
HVPRAVFVDLEPTVIDEVRTGTYRQLFHPEQLITGKEDAANNYARGHYTIGKEIIDLVLD
RIRKLADQCTGLQGFLVFHSFGGGTGSGFTSLLMERLSVDYGKKSKLEFSIYPAPQVSTA
VVEPYNSILTTHTTLEHSDCAFMVDNEAIYDICRRNLDIERPTYTNLNRLIGQIVSSITA
SLRFDGALNVDLTEFQTNLVPYPRIHFPLATYAPVISAEKAYHEQLSVAEITNACFEPAN
QMVKCDPRHGKYMACCLLYRGDVVPKDVNAAIATIKTKRTIQFVDWCPTGFKVGINYQPP
TVVPGGDLAKVQRAVCMLSNTTAIAEAWARLDHKFDLMYAKRAFVHWYVGEGMEEGEFSE
AREDMAALEKDYEEVGVDSVEGEGEEEGEEY

Ligands and cofactors

ID	Name	Formula	Copies
ADP	Adenosine-5'-diphosphate	C10 H15 N5 O10 P2	3
MG	Magnesium ion	Mg	4
GTP	Guanosine-5'-triphosphate	C10 H16 N5 O14 P3	5
ZN	Zinc ion	Zn	13

Primary citation

Structure of the mouse cytoplasmic lattice. Chi, P., Wang, X., Li, J. et al. Nature (2026) 654:523-531. DOI 10.1038/s41586-026-10442-6 · PubMed

Other PDB entries of the same protein (UniProt Q9D0L1 (AlphaFold model), which also has an AlphaFold model), best resolution first:

9L4J 3.0 Å, Structure of mouse SCMCcore complex contain ZBED3
24MC 3.4 Å, Structure of oocyte cytoplasmic lattices
12DL 3.5 Å, Native structure of the cytoplasmic lattice (CPL) asymmetric unit from mouse MII eggs
9SFP 4.2 Å, Native cytoplasmic lattices from mouse oocytes
9W2M 4.2 Å, Cryo-EM structure of the Cytoplasmic lattice(CPL) from mouse oocyte
9T63 4.7 Å, Asymmetric unit of a cytoplasmic lattice filament repeat unit
9T66 5.3 Å, Complex linking two repeat units of a cytoplasmic lattice filament
9T64 7.3 Å, Cytoplasmic lattice filament repeat unit with the central FBXW-SKP1 complex
9T67 7.7 Å, Cytoplasmic lattice filament repeat unit with ubiquitin
9T68 8.0 Å, Composite structure of the repeat unit of cytoplasmic lattice filament

Browse structure collections

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